Staff profile

1980 — 1981 — Postdoctoral Fellow (University of Oslo)

1981 – 1984  — Postdoctoral Fellow (University of Dundee)

1984 – 1998  — Lecturer — Reader in Palaeontology and Stratigraphy (University of Galway)

1998 – 2011  — Professor in Palaeontology (University of Copenhagen) 

2002 – 2011  — Deputy Head — Head of Geology (Natural History Museum of Denmark, University of Copenhagen) 

2004 – present  — Elected Foreign Member of the Royal Danish Academy of Sciences and Letters

2008 – present  — Einstein Professor, Chinese Academy of Sciences

2011 – 2021  — Professor of Palaeontology, Principal of Van Mildert College, Durham University

2013 – present  — Honorary Research Fellow, The Natural History Museum

2021 - present — Emeritus Professor of Palaeontology, Durham University

 

2011 – 2013  — Deputy Head of Colleges (Research and Scholarly Activities)

2012 – 2017 — Deputy Director, Climate Impacts Research Centre, Durham University

Career Research Highlights

My initial research on the stratigraphy and palaeontology of the Ordovician rocks of the Girvan district was largely monographic and was recognised by the Clough Memorial Award from the Edinburgh Geological Society. The description of Palaeozoic Brachiopoda using both conventional and statistical methodologies continues to be a focus of my research. The former led to an invitation to jointly revise with Sir Alwyn Williams various groups of Brachiopoda for the 'Treatise on Invertebrate Paleontology'; the latter has led to the development of microcomputer software - 'PALSTAT' package, which has now been completely rewritten and relaunched as 'PAST' in collaboration with Dr Øyvind Hammer (University of Oslo). The integration of such data into models of global biostratigraphy together with those for the environmental and spatial distributions of the phyla are continuing objectives within my research programme. This programme has been recognised with the award of the Lyell Fund from the Geological Society. Currently my main research interests have been modified regionally to include studies on the Lower Palaeozoic rocks in NE and N Greenland, Chile, China (including Tibet), Denmark, Estonia, Norway and Russia and my research in Greenland has been recognized by the award of Crown Prince Frederik's Fund. During the last ten years my focus has moved to target some of the larger scale processes in the history of life. Together with a range of colleagues, new models for biotic change and distributions through the Early Palaeozoic, particularly targeting the Cambrian Explosion, Great Ordovician Biodiversification Event and the end-Ordovician extinction, are being developed and their relationships to climatic and environmental changes are being assessed through a range of multidisciplinary techniques.

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Committee and Society Service

2018  — Member of Earth and Environmental Sciences panel, Portuguese Research Council

2015 — 2020 Member of advisory committee for School of Earth Space Sciences, Peking University

2015 — Member of evaluation panel, School of Earth Space Sciences, Peking University

2014 — Chair of evaluation panel, Paleontological Institute and Museum in the University of Zürich

2014 — Chair of evaluation panel, Earth Sciences programmes in the University of the West Indies

2014 —2016 President, Palaeontological Association

2012 – 2014  — Vice President, Palaeontographical Society

2012 – present  — NERC Peer Review College

2008 – present  — Voting member of the International Commission for Stratigraphy

2008 – present  — Chairman, International Subcommission on Ordovician Stratigraphy

2008 – 2009  — Member of evaluation panel of Earth Science Research Centres in Portugal

2007 – 2015  — Member of IGCP (UNESCO) scientific panel

2006 – 2010  — President, International Palaeontological Association

2005 – 2009; 2012  — Member of Earth Sciences panel, Swedish Research Council

2004 – 2009  — Co-leader IGCP (UNESCO) project 503

2004 – present  — Voting member of the International Ordovician Subcommission

2002 – 2004  — Vice President Palaeontological Association

1996 – 1998  — Vice President Irish Geological Association

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Conferences and Workshops (selected)

2018 — Co-convener ' IGCP 653' and 'Neogene tropical biotas and environments' sessions, 5th International Palaeontological Congress, Paris, France

2016 — Organiser, Opening Meeting of the 'Onset of the Great Ordovician Biodiversification Event'. International Geosciences Project 653, Van Mildert College, Durham University

2014 — Co-convener 'Ordovician biotas of Gondwana: responses to global climatic and eustatic events, and their biogeographic relationships within the Ordovician World’. 4^th International Palaeontological Congress, Mendoza, Argentina

2013 — Co-convener ‘The Cambrian Explosion – Understanding Earth systems at the origin of modern ecosystems’ Lyell Meeting, Geological Society, London

2012  — Co-convener 'International Subcommission on Ordovician stratigraphy: Ordovician intercontinental correlations: developing global and regional chronostratigraphy', International Geological Congress, Brisbane

2012  — Co-convener 'Out of Our Depth: The Paleontology, Ichnology, and Sedimentology of Deeper Water Environments in the Ancient Tropics', Annual Geological Society of America Meeting, Charlotte, North Carolina

2010  — Co-chair of science and vice-chair of executive committees for the 3rd International Palaeontological Congress, London

2009  — Organizer Final Meeting of IGCP 503, Copenhagen

2005  — Organizer 5th International Brachiopod Congress, Copenhagen

2003  — Co-convener 'Environmental and biological impact of the end Ordovician glaciation', EUG, Nice.

2002  — Co-organizer '4th Gilbert Larwood Symposium', Copenhagen

2001  — Organizer of the Annual Meeting of the Palaeontological Association, Copenhagen

2000  — Co-convener 'History of Biodiversity', Geosciences 2000, Manchester

2000  — Co-convener 'Biostratigraphy and Biogeography', Millennium Brachiopod Congress, London

2000  — Co-convener 'Tropics through Time', Annual Geological Society of America Meeting, Reno, Nevada

1998  — Co-organiser of Annual Irish Geological Research Meeting, University of Galway

1995  — Organiser of Annual Meeting of Palaeontological Association, University College, Galway

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Editorial Roles

2019 - present — Co editor-in-Chief, Palaeoworld

2017 - present — Associate Editor, Swiss Journal of Paleontology

2015 – 2019 — Member of Editorial Board, Nature Scientific Reports 

2014 - 2019 — Review Editor, Frontiers in Paleontology

2009 – 2017 — Advisory editor, Journal of the Geological Society, London

2008 – present  — Associate Editor, Paleontological Contributions

2005 – 2019 — Associate Editor, Palaeoworld

2004 – 2016  — Associate Editor, Swiss Journal of Geosciences

2004 – 2008  — Chair of Publications Board, Palaeontological Association

2003 – present  — Editor, Lethaia

1996 – 2012  — Associate Editor, Geological Journal

1996 – 2002  — Editor, for the Palaeontological Association's journal, Palaeontology

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Recent and Current Grants (Selected)

2006 – 2007  — Geocenter København (Geocenter Copenhagen): Mapping Denmark's and Greenland's palaeobiodiversity: An integrated database for Denmarks and Greenlands fossils. 400,000 DKK

2006  — Forskningsrådet for Natur og Univers (Danish Research Council): Rapid transitions from Greenhouse-Icehouse-Greenhouse climates 440 million year ago: New investigations of the palaeotropics in Early Palaeozoic North Greenland. 850,000 DKK

2006  — Forskningsrådet for Natur og Univers (Danish Research Council): Fossil microorganisms and ultrastructures in time and space (SEM equipment). 850,000 DKK

2006 – 2007  — INTERREG IIIA, Øresundsregionen: Integrated Higher Education in Geology (with Prof. Per Ahlberg, Lunds Universitet, Sverige). 500,000 SEK.

2007 – 2009  — Geocenter København (Geocenter Copenhagen): Monsters from deep time: Exhibition on Sirius Passet fossils. 450,000 DKK

2007  — TK-Foundation (Galathea III expedition). Ordovician in the roots of the Andes (with Prof. Lars Stemmerik). 300,000 DKK

2008 – 2012  — FNU (Danish Research Council) The Great Ordovician Biodiversification Event (GOBE). DKK 4,000,000 DKK

2009  — Geocenter Copenhagen: Expedition to Peary Land: Sirius Passet fauna and the Cambrian explosion. DKK 1,300,000

2010  — Carlsberg Foundation: Aspects of Modern and Ancient diversity on the Tibetan Plateau. 500,000 DKK

2011  — Carlsberg Foundation: The Dawn of Phanerozoic ecosystems. DKK 500,000

2011  — Agouron Institute: The Cambrian Explosion in the Sirius Pass, North Greenland. DKK 900,000. (co applicant with Prof. Don Canfield).

2012 – 2014  — FNU (Danish Research Council): Unravelling early animal evolution. DKK 2,000,000

2016 – SEK Wenner-Gren Foundation: Cambrian brachiopods ‒ diversity, distribution and evolution. SEK 106, 000 (co applicant with Prof. Per Ahlberg)

2016 – 2018 – Leverhulme Trust: Roots of the Great Ordovician Biodiversification. £49,000

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Research Interests

• Fossil brachiopods, their taxonomy, distribution, ecology and phylogeny.
• Major events in the history life, e.g. Cambrian Explosion, Great Ordovician Biodiversification Event and end Ordovician extinction
• Numerical methods in palaeontology

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• 2019: Research Associate; National Museum of Scotland:
• 2019: Guest Research Professor, Chinese Academy of Sciences, Nanjing:
• 2017: Distinguished Visiting Professor, China University of Geosciences, Wuhan:
• 2016: Leverhulme Fellow:
• 2016: Chair, International Commission on Stratigraphy:
• 2014: President, Palaeontological Association:
• 2014: Visiting Professor, CNRS - Université de Lille 1, France:
• 2014: Guest Professor, University of Lund:
• 2014: Elected Foreign Member of the Royal Swedish Physiographic Society:
• 2014: Member of Scientific Advisory Board, Naturalis (State Museum of Natural History), The Netherlands:
• 2014: William King Medal, National University of Ireland, Galway:
• 2013: Honorary Research Fellow, The Natural History Museum:
• 2008: Einstein Professor, Chinese Academy of Sciences:
• 2004: Elected Foreign Member of the Royal Danish Academy of Sciences and Letters:
• 2003: Crown Prince Frederik's Fund, Denmark:
• 1996: Lyell Fund from the Geological Society:
• 1986: Clough Memorial Award from the Edinburgh Geological Society:

Authored book

• Benton, M., & Harper, D. (2020). Introduction to Paleobiology and the Fossil Record, 2nd edition. Wiley
• Liljeroth, M., Harper, D., Carlisle, H., & Nielsen, A. (2017). Ordovician rhynchonelliformean brachiopods from Co. Waterford, SE Ireland: Palaeobiogeography of the Leinster Terrane. John Wiley and Sons. https://doi.org/10.1002/9781119412595
• Jiayu, R., Bing, H., Ren-bin, Z., & Harper, D. (2013). Latest Ordovician and earliest Silurian brachiopods succeeding the Hirnantia fauna in south-east China. Palaeontological Association. https://doi.org/10.1111/pala.12056
• Rasmussen, C., Harper, D., & Blodgett, R. (2012). Late Ordovician Brachiopods from West-Central Alaska: systematics, ecology and palaeobiogeography. Wiley
• Benton, M., & Harper, D. (2009). Introduction to Paleobiology and the Fossil Record. John Wiley and Sons

Chapter in book

• Harper, D. A. (2021). Brachiopoda. In D. Alderton, & S. A. Elias (Eds.), Encyclopedia of Geology (273-282). (2nd). Elsevier. https://doi.org/10.1016/b978-0-08-102908-4.00108-9
• Harper, D. A. (2021). Late Ordovician Extinctions. In D. Alderton, & S. A. Elias (Eds.), Encyclopedia of Geology (617-626). (2nd). Elsevier. https://doi.org/10.1016/b978-0-12-409548-9.12530-8
• Harper, D. A. (2021). Paleontology. In J. Vonk, & T. Shackelford (Eds.), Encyclopedia of Animal Cognition and Behavior. Springer Verlag. https://doi.org/10.1007/978-3-319-47829-6_527-1
• Armstrong, H., & Harper, D. (2014). An earth system approach to understanding the end-Ordovician (Hirnantian) mass extinction. In G. Keller, & A. Kerr (Eds.), Volcanism, impacts, and mass extinctions : causes and effects (287-300). Geological Society of America. https://doi.org/10.1130/2014.2505%2814%29
• Platform: North Atlantic Region. In J. Derby, R. Fritz, S. Longacre, W. Morgan, & C. Sternbach (Eds.), The great American carbonate bank: The geology and economic resources of the Cambrian-Ordovician Sauk megasequence of Laurentia (597-626). AAPG Memoir

Edited book

• Harper, D., & Servais, T. (Eds.). (2013). Early Palaeozoic Biogeography and Palaeogeography. Geological Society

Journal Article

• Murray, J., Nasheuer, H., Seoighe, C., McCormick, G., Williams, D., & Harper, D. (in press). The contribution of William King to the early development of palaeoanthropology. Irish Journal of Earth Sciences, 33, 1-16. https://doi.org/10.3318/ijes.2015.33.1
• Penn-Clarke, C. R., & Harper, D. A. (in press). Early−Middle Devonian brachiopod provincialism and bioregionalization at high latitudes: A case study from southwestern Gondwana. GSA Bulletin, https://doi.org/10.1130/b35670.1
• Harper, D., Zhan, R., & Jin, J. (in press). The Great Ordovician Biodiversification Event: Reviewing two decades of research on diversity’s big bang illustrated by mainly brachiopod data. Palaeoworld, 24(1-2), 75-85. https://doi.org/10.1016/j.palwor.2015.03.003
• Guo, Z., Chen, Z., & Harper, D. A. (in press). Phylogenetic and ecomorphologic diversifications of spiriferinid brachiopods after the end-Permian extinction. Paleobiology, 1-16. https://doi.org/10.1017/pab.2020.34
• Mussini, G., Smith, M. P., Vinther, J., Rahman, I. A., Murdock, D. J. E., Harper, D. A. T., & Dunn, F. S. (2024). A new interpretation of Pikaia reveals the origins of the chordate body plan. Current Biology, Article S0960-9822(24)00669-9. https://doi.org/10.1016/j.cub.2024.05.026
• Jin, J., & Harper, D. A. (2024). An Edgewood-type Hirnantian fauna from the Mackenzie Mountains, northwestern margin of Laurentia. Journal of Paleontology, 98(1), 13-39. https://doi.org/10.1017/jpa.2023.87
• Jin, J., Rasmussen, C. M. Ø., Sheehan, P. M., & Harper, D. A. T. (2024). Late Ordovician and early Silurian virgianid and stricklandioid brachiopods from North Greenland: implications for a warm‐water faunal province. Papers in Palaeontology, 10(1), Article e1544. https://doi.org/10.1002/spp2.1544
• Harper, D. A. T. (2024). Late Ordovician Mass Extinction: Earth, fire and ice. National Science Review, 11(1), Article nwad319. https://doi.org/10.1093/nsr/nwad319
• Park, T. S., Nielsen, M. L., Parry, L. A., Sørensen, M. V., Lee, M., Kihm, J., …Vinther, J. (2024). A giant stem-group chaetognath. Science Advances, 10(1), Article eadi6678. https://doi.org/10.1126/sciadv.adi6678
• Wu, S., Reitner, J., Harper, D. A. T., Yu, J., & Chen, Z.-Q. (2023). New keratose sponges after the end-Permian extinction provide insights into biotic recoveries. Geobiology, 22(1), Article e12582. https://doi.org/10.1111/gbi.12582
• Harper, D. A., & Bates, D. E. (2023). Middle Ordovician brachiopods from Tagoat, Co. Wexford, SE Ireland: Dapingian diversity drivers. Geobios, 81, 85-100. https://doi.org/10.1016/j.geobios.2023.06.006
• Penn-Clarke, C. R., & Harper, D. A. (2023). The rise and fall of the Malvinoxhosan (Malvinokaffric) bioregion in South Africa: Evidence for Early-Middle Devonian biocrises at the South Pole. Earth-Science Reviews, 246, Article 104595. https://doi.org/10.1016/j.earscirev.2023.104595
• Servais, T., Cascales-Miñana, B., Harper, D. A., Lefebvre, B., Munnecke, A., Wang, W., & Zhang, Y. (2023). No (Cambrian) explosion and no (Ordovician) event: A single long-term radiation in the early Palaeozoic. Palaeogeography, Palaeoclimatology, Palaeoecology, 623, Article 111592. https://doi.org/10.1016/j.palaeo.2023.111592
• Harper, D. A., Meidla, T., & Servais, T. (2023). A short history of the Ordovician System: from overlapping unit stratotypes to global stratotype sections and points. Geological Society Special Publications, 532(1), https://doi.org/10.1144/sp532-2022-285
• Murray, J., MacGabhann, B. A., Doyle, E., Mángano, M. G., Tyrrell, S., & Harper, D. A. (2023). An enigmatic large discoidal fossil from the Pennsylvanian of County Clare, Ireland. Palaeoworld, https://doi.org/10.1016/j.palwor.2023.01.008
• Harper, D. A. (2023). The Irish Ordovician brachiopod fauna: A taxonomic renaissance. Estonian Journal of Earth Sciences, 72(1), 38-41. https://doi.org/10.3176/earth.2023.35
• Hughes, N. C., Peng, S., Harper, D. A., Myrow, P. M., Phạm, N. K., Wernette, S. J., & Zhu, X. (2022). Cambrian and earliest Ordovician fauna and geology of the Sông Đà and adjacent terranes in Việt Nam (Vietnam). Geological Magazine, 159(1), 55-80. https://doi.org/10.1017/s0016756821000844
• Saleh, F., Vaucher, R., Vidal, M., Hariri, K. E., Laibl, L., Daley, A. C., …Lefebvre, B. (2022). New fossil assemblages from the Early Ordovician Fezouata Biota. Scientific Reports, 12(1), Article 20773. https://doi.org/10.1038/s41598-022-25000-z
• Aubry, M., Piller, W. E., Gibbard, P. L., Harper, D. A., & Finney, S. C. (2022). Ratification of subseries/subepochs as formal rank/units in international chronostratigraphy. Episodes, 45(1), 97-99. https://doi.org/10.18814/epiiugs/2021/021016
• Pereira, S., Colmenar, J., Mortier, J., Vanmeirhaeghe, J., Verniers, J., Štorch, P., …Gutiérrez-Marco, J. C. (2021). Hirnantia Fauna from the Condroz Inlier, Belgium: another case of a relict Ordovician shelly fauna in the Silurian?. Journal of Paleontology, 6, 1189-1215. https://doi.org/10.1017/jpa.2021.74
• Zhang, M., Chen, Z., & Harper, D. A. (2021). An atypical Burgess Shale-type fossil assemblage from Cambrian Stage 4 of the Jingshan area, South China: Taphonomy, palaeoecology, and global correlations. Global and Planetary Change, 206, Article 103640. https://doi.org/10.1016/j.gloplacha.2021.103640
• Simonet Roda, M., Griesshaber, E., Angiolini, L., Harper, D. A., Jansen, U., Bitner, M. A., …Schmahl, W. W. (2021). The evolution of thecideide microstructures and textures: traced from Triassic to Holocene. Lethaia: An International Journal of Palaeontology and Stratigraphy, 54(4), 558-577. https://doi.org/10.1111/let.12422
• Harper, D. A., Stouge, S., Christiansen, J. L., Topper, T. P., Alwmark, C., Richoz, S., & Ahlberg, P. (2021). Early Cambrian brachiopod-dominated shell concentrations from North-East Greenland: Environmental and taphonomic implications. Global and Planetary Change, 204, Article 103560. https://doi.org/10.1016/j.gloplacha.2021.103560
• Donovan, S. K., Jones, S. J., King, A. R., & Harper, D. A. (2021). Pliocene trace fossils from oyster substrates in the Nijar Basin, Betic Cordillera, southern Spain. Proceedings of the Geologists' Association, 132(3), 358-368. https://doi.org/10.1016/j.pgeola.2021.02.004
• Servais, T., Cascales-Minana, B., & Harper, D. (2021). The Great Ordovician Biodiversification Event (GOBE) is not a single event. Paleontological Research, 25(4), 315-328. https://doi.org/10.2517/2021pr001
• Guo, Z., Chen, Z., Harper, D. A., & Huang, Y. (2021). Permian–Triassic phylogenetic and morphologic evolution of rhynchonellide brachiopods. Paleobiology, https://doi.org/10.1017/pab.2021.25
• Harper, D. A., Cascales-Miñana, B., Kroeck, D. M., & Servais, T. (2021). The palaeogeographical impact on the biodiversity of marine faunas during the Ordovician radiations. Global and Planetary Change, 207, Article 103665. https://doi.org/10.1016/j.gloplacha.2021.103665
• Jiayu, R., Harper, D., Bing, H., Rongyu, L., Xiaole, Z., & Di, C. (2020). The latest Ordovician Hirnantian brachiopod faunas: New global insights. Earth-Science Reviews, 208, Article 103280. https://doi.org/10.1016/j.earscirev.2020.103280
• Zhen, Y. Y., Zhang, Y., Harper, D. A., Zhan, R., Fang, X., Wang, Z., …Li, W. (2020). Ordovician successions in southern-central Xizang (Tibet), China—Refining the stratigraphy of the Himalayan and Lhasa terranes. Gondwana Research, 83, 372-389. https://doi.org/10.1016/j.gr.2020.01.023
• Rong, J., Aung, K. P., Zhan, R., Huang, B., Harper, D. A., Chen, D., …Zhang, X. (2020). The latest Ordovician Hirnantia brachiopod fauna of Myanmar: Significance of new data from the Mandalay Region. Palaeoworld, 29(1), 1-30. https://doi.org/10.1016/j.palwor.2019.07.002
• Huang, B., Rong, J., Harper, D. A., & Zhou, H. (2020). A nearshore Hirnantian brachiopod fauna from South China and its ecological significance. Journal of Paleontology, 94(2), 239-254. https://doi.org/10.1017/jpa.2019.90
• Huang, B., Zhou, H., Harper, D. A., Zhan, R., Zhang, X., Chen, D., & Rong, J. (2020). A latest Ordovician Hirnantia brachiopod fauna from western Yunnan, Southwest China and its paleobiogeographic significance. Palaeoworld, 29(1), 31-46. https://doi.org/10.1016/j.palwor.2019.03.002
• Harper, D. A., Cascales-Miñana, B., & Servais, T. (2020). Early Palaeozoic diversifications and extinctions in the marine biosphere: a continuum of change. Geological Magazine, 157, 5-21. https://doi.org/10.1017/s0016756819001298
• curiosity. Irish Journal of Earth Sciences, 38, 73-90. https://doi.org/10.3318/ijes.2020.38.5
• Stouge, S., Harper, D. A., Zhan, R., Liu, J., & Stemmerik, L. (2020). Middle Ordovician (Darriwilian) conodonts from southern Tibet, the Indian passive margin: implications for the age and correlation of the roof of the world. Geological Magazine, 1-25. https://doi.org/10.1017/s0016756820001077
• Guo, Z., Chen, Z., & Harper, D. A. (2020). The Anisian (Middle Triassic) brachiopod fauna from Qingyan, Guizhou, south-western China. Journal of Systematic Palaeontology, 18(8), 647-701. https://doi.org/10.1080/14772019.2019.1682695
• Rønning, K. J., Bruton, D. L., Harper, D. A., Høyberget, M., Maletz, J., & Nakrem, H. A. (2020). A Cambrian–Ordovician boundary section in the Rafnes–Herøya submarine tunnel, Skien–Langesund District, southern Norway. Norsk Geologisk Tidsskrift, 100(1), 1-24. https://doi.org/10.17850/njg100-3-3
• Ling, M., Zhan, R., Wang, G., Wang, Y., Amelin, Y., Tang, P., …Rong, J. (2019). An extremely brief end Ordovician mass extinction linked to abrupt onset of glaciation. Solid earth sciences, 4(4), 190-198. https://doi.org/10.1016/j.sesci.2019.11.001
• Saleh, F., Candela, Y., Harper, D. A., Polechová, M., Lefebvre, B., & Pittet, B. (2019). Storm-induced community dynamics in the Fezouata Biota (Lower Ordovician, Morocco). PALAIOS, 33(12), 535-541. https://doi.org/10.2110/palo.2018.055
• Harper, D., Hammarlund, E., Topper, T., Nielsen, A., Rasmussen, J., T-y.s., P., & Smith, M. (2019). The Sirius Passet Lagerstätte of North Greenland: a remote window on the Cambrian Explosion. Journal of the Geological Society, 176(6), 1023-1037. https://doi.org/10.1144/jgs2019-043
• Servais, T., Cascales-Miñana, B., Cleal, C. J., Gerrienne, P., Harper, D. A., & Neumann, M. (2019). Revisiting the Great Ordovician Diversification of land plants: Recent data and perspectives. Palaeogeography, Palaeoclimatology, Palaeoecology, 534, https://doi.org/10.1016/j.palaeo.2019.109280
• Angiolini, L., Crippa, G., Azmy, K., Capitani, G., Confalonieri, G., Della Porta, G., …Stephenson, M. H. (2019). The giants of the phylum Brachiopoda: a matter of diet?. Palaeontology, 62(6), 889-917. https://doi.org/10.1111/pala.12433
• Schmitz, B., Farley, K. A., Goderis, S., Heck, P. R., Bergström, S. M., Boschi, S., …Terfelt, F. (2019). An extraterrestrial trigger for the mid-Ordovician ice age: Dust from the breakup of the L-chondrite parent body. Science Advances, 5(9), Article eaax4184. https://doi.org/10.1126/sciadv.aax4184
• Huang, B., Harper, D. A., Zhou, H., Zhan, R., Wang, Y., Tang, P., …Rong, J. (2019). A new Cathaysiorthis (Brachiopoda) fauna from the lower Llandovery of eastern Qinling, China. Papers in Palaeontology, 5(3), 537-557. https://doi.org/10.1002/spp2.1253
• Harper, D. (2019). The Golden Spike Still Glitters: The (Re)construction of a Global Chronostratigraphy. Acta Geologica Sinica - English Edition, 93(S3), 24-27. https://doi.org/10.1111/1755-6724.14234
• Harper, D. A., Topper, T. P., Cascales-Miñana, B., Servais, T., Zhang, Y., & Ahlberg, P. (2019). The Furongian (late Cambrian) Biodiversity Gap: Real or apparent?. Palaeoworld, 28(1-2), 4-12. https://doi.org/10.1016/j.palwor.2019.01.007
• Hammarlund, E. U., Smith, M. P., Rasmussen, J. A., Nielsen, A. T., Canfield, D. E., & Harper, D. A. (2019). The Sirius Passet Lagerstätte of North Greenland—A geochemical window on early Cambrian low‐oxygen environments and ecosystems. Geobiology, 17(1), 12-26. https://doi.org/10.1111/gbi.12315
• Huang, B., Harper, D. A., Zhou, H., & Rong, J. (2019). From shallow to deep water: an ecological study of the Hirnantia brachiopod Fauna (Late Ordovician) and its global implications. Lethaia: An International Journal of Palaeontology and Stratigraphy, 1-13. https://doi.org/10.1111/let.12360
• Drost, K., Chew, D., Petrus, J. A., Scholze, F., Woodhead, J. D., Schneider, J. W., & Harper, D. A. (2018). An Image Mapping Approach to U-Pb LA-ICP-MS Carbonate Dating and Applications to Direct Dating of Carbonate Sedimentation. Geochemistry, Geophysics, Geosystems, 19(12), 4631-4648. https://doi.org/10.1029/2018gc007850
• Bogolepova, O. K., Donovan, S. K., Harper, D. A., Suyarkova, A. A., Yakupov, R., & Gubanov, A. P. (2018). New records of brachiopods and crinoids from the Silurian (Wenlock) of the southern Urals, Russia. GFF, 140(4), 323-331. https://doi.org/10.1080/11035897.2018.1526210
• Hints, L., Harper, D., & Paškevičius, J. (2018). Diversity and biostratigraphic utility of Ordovician brachiopods in the East Baltic. Estonian Journal of Earth Sciences, 67(3), 176-191. https://doi.org/10.3176/earth.2018.14
• Strang, K., Harper, D., Donovan, S., & Portell, R. (2018). Silicification of low-magnesium mollusc shells from the Upper Oligocene of Antigua, Lesser Antilles
• Harper, D., & Servais, T. (2018). Contextualizing the Onset of the Great Ordovician Biodiversification Event. Lethaia: An International Journal of Palaeontology and Stratigraphy, 51(2), 149-150. https://doi.org/10.1111/let.12264
• Topper, T. P., Greco, F., Hofmann, A., Beeby, A., & Harper, D. A. (2018). Characterization of kerogenous films and taphonomic modes of the Sirius Passet Lagerstätte, Greenland. Geology, 46(4), 359-362. https://doi.org/10.1130/g39930.1
• Topper, T., Zhang, Z., Gutiérrez-Marco, J., & Harper, D. (2018). The dawn of a dynasty: life strategies of Cambrian and Ordovician brachiopods. Lethaia: An International Journal of Palaeontology and Stratigraphy, 51(2), 254-266. https://doi.org/10.1111/let.12229
• Pohl, A., Harper, D. A., Donnadieu, Y., Le Hir, G., Nardin, E., & Servais, T. (2018). Possible patterns of marine primary productivity during the Great Ordovician Biodiversification Event. Lethaia: An International Journal of Palaeontology and Stratigraphy, 51(2), 187-197. https://doi.org/10.1111/let.12247
• Servais, T., & Harper, D. A. (2018). The Great Ordovician Biodiversification Event (GOBE): definition, concept and duration. Lethaia: An International Journal of Palaeontology and Stratigraphy, 51(2), 151-164. https://doi.org/10.1111/let.12259
• Park, T. S., Kihm, J., Woo, J., Park, C., Lee, W. Y., Smith, M. P., …Vinther, J. (2018). Brain and eyes of Kerygmachela reveal protocerebral ancestry of the panarthropod head. Nature Communications, 9(1), Article 1019. https://doi.org/10.1038/s41467-018-03464-w
• Erlykin, A. D., Harper, D. A., Sloan, T., & Wolfendale, A. W. (2018). Periodicity in extinction rates. Palaeontology, 61(1), 149-158. https://doi.org/10.1111/pala.12334
• Blake, D. B., Donovan, S. K., & Harper, D. A. (2017). A new Silurian ophiuroid from the west of Ireland. Irish Journal of Earth Sciences, 35, 57-66. https://doi.org/10.3318/ijes.2017.35.57
• Finnegan, S., Rasmussen, C., & Harper, D. (2017). Identifying the most surprising victims of mass extinction events: an example using Late Ordovician Brachiopods. Biology Letters, 13(9), Article 2017400. https://doi.org/10.1098/rsbl.2017.0400
• Nielsen, M., Rasmussen, J., & Harper, D. (2017). Sexual dimorphism within the stem-group arthropod Isoxys volucris from the Sirius Passet Lagerstätte, North Greenland. Bulletin of the Geological Society of Denmark, 65, 47-58
• Harper, D. A., Popov, L. E., & Holmer, L. E. (2017). Brachiopods: origin and early history. Palaeontology, 60(5), 609-631. https://doi.org/10.1111/pala.12307
• Harper, D. A., Parkes, M. A., & Ren-Bin, Z. (2017). Late Ordovician deep-water brachiopod fauna from Raheen, Waterford Harbour, Ireland. Irish Journal of Earth Sciences, 35, 1-18. https://doi.org/10.3318/ijes.2017.35.1
• Donovan, S. K., Harper, D. A., Portell, R. W., & Toomey, J. K. (2017). Echinoids as hard substrates: varied examples from the Oligocene of Antigua, Lesser Antilles. Proceedings of the Geologists' Association, 128(3), 326-331. https://doi.org/10.1016/j.pgeola.2017.04.003
• Huang, B., Harper, D. A., Rong, J., & Zhan, R. (2017). Brachiopod faunas after the end Ordovician mass extinction from South China: Testing ecological change through a major taxonomic crisis. Journal of Asian Earth Sciences, 138, 502-514. https://doi.org/10.1016/j.jseaes.2017.02.043
• Erlykin, A. D., Harper, D. A., Sloan, T., & Wolfendale, A. W. (2017). Mass extinctions over the last 500 myr: an astronomical cause?. Palaeontology, 60(2), 159-167. https://doi.org/10.1111/pala.12283
• Donovan, S., Harper, D., & Portell, R. (2017). Shell-Filled Burrows in the Upper Oligocene Antigua Formation, Antigua, Lesser Antilles. Ichnos: an International Journal of Plant and Animal Traces, 24, 72-77. https://doi.org/10.1080/10420940.2016.1223653
• Strang, K., Armstrong, H., & Harper, D. (2016). Minerals in the gut: scoping a Cambrian digestive system. Royal Society Open Science, 3(11), Article 160420. https://doi.org/10.1098/rsos.160420
• Donovan, S., Jones, B., & Harper, D. (2016). Neogene echinoids from the Cayman Islands, West Indies: regional implications. Geological Journal, 51(6), 864-879. https://doi.org/10.1002/gj.2703
• Strang, K., Armstrong, H., Harper, D., & Trabucho-Alexandre, J. P. (2016). The Sirius Passet Lagerstatte: silica death masking opens the window on the earliest matground community of the Cambrian explosion. Lethaia: An International Journal of Palaeontology and Stratigraphy, 49(4), 631-643. https://doi.org/10.1111/let.12174
• Chenyi, T., Zhong-Qiang, C., & Harper, D. (2016). Permian–Triassic evolution of the Bivalvia: Extinction-recovery patterns linked to ecologic and taxonomic selectivity. Palaeogeography, Palaeoclimatology, Palaeoecology, 459, 53-62. https://doi.org/10.1016/j.palaeo.2016.06.042
• Jensen, S., Harper, D., & Stouge, S. (2016). Trace fossils from the lower Cambrian Kløftelv Formation, Ella Ø, North-East Greenland. GFF, 138(3), 369-376. https://doi.org/10.1080/11035897.2015.1076029
• their palaeogeographic implication. Geological Journal, 51(4), 584-599. https://doi.org/10.1002/gj.2659
• Jakobsen, K., Brock, G., Nielsen, A., & Harper, D. (2016). A Darriwilian (Middle Ordovician) bivalve-dominated molluscan fauna from the Stairway Sandstone, Amadeus Basin, central Australia. Acta Palaeontologica Polonica, 61(4), 897-924. https://doi.org/10.4202/app.00215.2015
• Finnegan, S., Rasmussen, C., & Harper, D. (2016). Biogeographic and bathymetric determinants of brachiopod extinction and survival during the Late Ordovician mass extinction. Proceedings of the Royal Society B: Biological Sciences, 283(1829), Article 20160007. https://doi.org/10.1098/rspb.2016.0007
• Harper, D., & Hints, L. (2016). Hirnantian (Late Ordovician) brachiopod faunas across Baltoscandia: A global and regional context. Palaeogeography, Palaeoclimatology, Palaeoecology, 444, 71-83. https://doi.org/10.1016/j.palaeo.2015.11.044
• Rasmussen, C. Ø., Ullmann, C., Jakobsen, K., Lindskog, A., Hansen, J., Hansen, T., …Harper, D. (2016). Onset of main Phanerozoic marine radiation sparked by emerging Mid Ordovician icehouse. Scientific Reports, 6, Article 18884. https://doi.org/10.1038/srep18884
• Rohr, D., Harper, D., Stouge, S., & Christiansen, J. (2015). Ordovician Gastropoda from Northeast Greenland. Bulletin of Geosciences (On-line), 90, 795-805. https://doi.org/10.3140/bull.geosci.1566
• Donovan, S., Harper, D., & Portell, R. (2015). In deep water: a crinoid–brachiopod association in the Upper Oligocene of Antigua, West Indies. Lethaia: An International Journal of Palaeontology and Stratigraphy, 48(3), 291-298. https://doi.org/10.1111/let.12105
• Zhan, R., Jin, J., & Harper, D. (2015). Ecosystem revolution and evolution in the Early–Mid Paleozoic. Palaeoworld, 24(1-2), 1-4. https://doi.org/10.1016/j.palwor.2015.05.001
• Jakobsen, K., Brock, G., Nielsen, A., Mathieson, D., & Harper, D. (2015). Brachiopods associated with stromatoporoid mounds from the Middle to Upper Ordovician Cashions Creek Limestone, Tasmania. Memoir ... of the Association of Australasian Palaeontologists, 45, 249-265
• Hints, L., & Harper, D. (2015). The Hirnantian (Late Ordovician) brachiopod fauna of the East Baltic: Taxonomy of the key species. Acta Palaeontologica Polonica, 60(2), 395-420. https://doi.org/10.4202/app.2013.0010
• the Upper Ordovician of North America. Papers in Palaeontology, 1, 237-253. https://doi.org/10.1002/spp2.1014
• Jagt, J., Thuy, B., Donovan, S., Stohr, S., Portell, R., Pickerill, R., …Jackson, T. (2014). A starfish bed in the Middle Miocene Grand Bay Formation of Carriacou, The Grenadines (West Indies). Geological Magazine, 151(3), 381-393. https://doi.org/10.1017/s0016756813000204
• Harper, D., Hammarlund, E., & Rasmussen, C. (2014). End Ordovician extinctions : a coincidence of causes. Gondwana Research, 25(4), 1294-1307. https://doi.org/10.1016/j.gr.2012.12.021
• Vinther, J., Stein, M., Longrich, N., & Harper, D. (2014). A suspension-feeding anomalocarid from the Early Cambrian. Nature, 507(7493), 496-499. https://doi.org/10.1038/nature13010
• Renbin, Z., Harper, D., Jisuo, J., Yan, L., Jianbo, L., Stemmerik, L., & Stouge, S. (2014). Middle Ordovician Aporthophyla brachiopod fauna from the roof of the World, Southern Tibet. Palaeontology, 57(1), 141-170. https://doi.org/10.1111/pala.12058
• Bing, H., Harper, D., & Renbin, Z. (2014). Test of sampling sufficiency in palaeontology. GFF, 136(1), 105-109. https://doi.org/10.1080/11035897.2014.882976
• Colmenar, J., Harper, D., & Villas, E. (2014). Morphofunctional analysis of Svobodaina species (Brachiopoda, Heterorthidae) from South-Western Europe. Palaeontology, 57(1), 193-214. https://doi.org/10.1111/pala.12061
• Harper, D., Jisuo, J., & Rasmussen, C. (2014). Late Ordovician carbonate mounds from North Greenland: a peri-Laurentian dimension to the Boda Event?. GFF, 136(1), 95-99. https://doi.org/10.1080/11035897.2013.865669
• Servais, T., Danielian, T., Harper, D., & Munnecke, A. (2014). Possible oceanic circulation patterns, surface water currents and upwelling zones in the Early Palaeozoic. GFF, 136(1), 229-233. https://doi.org/10.1080/11035897.2013.876659
• Jakobsen, K., Nielsen, A., Harper, D., & Brock, G. (2014). Trilobites from the Middle Ordovician Stairway Sandstone, Amadeus Basin, central Australia. Alcheringa: An Australasian Journal of Palaeontology, 38, 70-96. https://doi.org/10.1080/03115518.2013.828253
• Oligocene oysters, Antigua, West Indies. Proceedings of the Geologists' Association, 125(1), 99-106. https://doi.org/10.1016/j.pgeola.2013.10.002
• Smith, M., & Harper, D. (2013). Causes of the Cambrian Explosion. Science, 341(6152), 1355-1356. https://doi.org/10.1126/science.1239450
• Stein, M., Budd, G., Peel, J., & Harper, D. (2013). Arthroaspis n. gen., a common element of the Sirius Passet Lagerstätte (Cambrian, North Greenland), sheds light on trilobite ancestry. BMC Evolutionary Biology, 13, Article 99. https://doi.org/10.1186/1471-2148-13-99
• Servais, T., Cecca, F., Harper, D., Isozaki, Y., & Mac Niocaill, C. (2013). Palaeozoic palaeogeographical and palaeobiogeographical nomenclature. Memoirs, 38(1), 25-33. https://doi.org/10.1144/m38.3
• Topper, T., Skovsted, C., Peel, J., & Harper, D. (2013). Moulting in the lobopodian Onychodictyon from the lower Cambrian of Greenland. Lethaia: An International Journal of Palaeontology and Stratigraphy, 46, 490-495. https://doi.org/10.1111/let.12026
• Zuykov, M., Pelletier, E., & Harper, D. (2013). Bivalve mollusks in metal pollution studies: from bioaccumulation to biomonitoring. Chemosphere, 93, 201-208. https://doi.org/10.1016/j.chemosphere.2013.05.001
• Topper, T., Harper, D., & Brock, G. (2013). Ancestral billingsellides and the evolution and phylogenetic relationships of early rhynchonelliform brachiopods. Journal of Systematic Palaeontology, 11(7), 821-833. https://doi.org/10.1080/14772019.2012.728253
• Jakobsen, K., Brock, G., Nielsen, A., Topper, T., & Harper, D. (2013). Middle Ordovician brachiopods from the Stairway Sandstone, Amadeus Basin, central Australia. Alcheringa: An Australasian Journal of Palaeontology, 38, 190-208. https://doi.org/10.1080/03115518.2013.858204
• Bing, H., Harper, D., & Hammer, O. (2013). Introduction to PAST, a comprehensive statistics software package for paleontological data analysis. Gushengwu xuebao, 52, 161-181
• rhynchonelliform brachiopods. Memoirs, 38(1), 127-144. https://doi.org/10.1144/m38.11
• Tvären crater system, Sweden. Palaeogeography, Palaeoclimatology, Palaeoecology, 369, 114-124. https://doi.org/10.1016/j.palaeo.2012.10.008
• Jisuo, J., Harper, D., Cocks, L., McCausland, P., Rasmussen, C., & Sheehan, P. (2013). Precisely locating the Ordovician equator in Laurentia. Geology, 41(2), 107-110. https://doi.org/10.1130/g33688.1
• 150 years of confusion. GFF, 135(2), 191-203. https://doi.org/10.1080/11035897.2013.811440
• Harper, D., & Servais, T. (2013). Early Palaeozoic biogeography and palaeogeography: towards a modern synthesis. Memoirs, 38(1), 1-3. https://doi.org/10.1144/m38.1
• Topper, T., Skovsted, C., Harper, D., & Ahlberg, P. (2013). A Bradoriid and Brachiopod Dominated Shelly Fauna from the Furongian(Cambrian) of Västergötland, Sweden. Journal of Paleontology, 87(1), 69-83. https://doi.org/10.1666/12-047r.1
• Mangano, M., Bromley, R., Harper, D., Nielsen, A., Smith, M., & Vinther, J. (2012). Nonbiomineralized carapaces in Cambrian seafloor landscapes (Sirius Passet, Greenland): Opening a new window into early Phanerozoic benthic ecology. Geology, 40(6), 519-522. https://doi.org/10.1130/g32853.1
• Hammarlund, E., Dahl, T., Harper, D., Bond, D., Nielsen, A., Bjerrum, C., …Canfield, D. (2012). A sulfidic driver for the end-Ordovician mass extinction. Earth and Planetary Science Letters, 331-332, 128-139. https://doi.org/10.1016/j.epsl.2012.02.024
• of Laurentia. Palaeogeography, Palaeoclimatology, Palaeoecology, 367-368, 73-88. https://doi.org/10.1016/j.palaeo.2011.05.023
• Donovan, S., Harper, D., Renbin, Z., Stemmerik, L., Jianbo, L., & Stouge, S. (2012). A primitive cladid crinoid from the Jiacun Group, Tibet (Darriwilian, Middle Ordovician). Geological Journal, 47, 653-660. https://doi.org/10.1002/gj.2436
• Zuykov, M., Vinarski, M., Pelletier, E., Demers, S., & Harper, D. (2012). Shell malformations in seven species of pond snail (Gastropoda, Lymnaeidae): analysis of large museum collections. Zoosystematics and Evolution, 88(2), 365-368. https://doi.org/10.1002/zoos.201200025
• Topper, T., Holmer, L., Skovsted, C., Brock, G., Balthasar, U., Larsson, C., …Harper, D. (2012). The oldest brachiopods from the lower Cambrian of South Australia. Acta Palaeontologica Polonica, 58(1), 93-109. https://doi.org/10.4202/app.2011.0146
• Rasmussen, C., & Harper, D. (2011). Did the amalgamation of continents drive the end Ordovician mass extinctions?. Palaeogeography, Palaeoclimatology, Palaeoecology, 311(1-2), 48-62. https://doi.org/10.1016/j.palaeo.2011.07.029
• Rasmussen, C., & Harper, D. (2011). Interrogation of distributional data for the End Ordovician crisis interval: where did disaster strike?. Geological Journal, 46, 478-500. https://doi.org/10.1002/gj.1310
• Vinther, J., Eibye-Jacobsen, D., & Harper, D. (2011). An Early Cambrian stem polychaete with pygidial cirri. Biology Letters, 7(6), 929-932. https://doi.org/10.1098/rsbl.2011.0592
• Stouge, S., Christiansen, J., Harper, D., Houmark-Nielsen, M., Kristiansen, K., MacNiocaill, C., & Buchard-Westergaard, B. (2011). Neoproterozoic (Cryogenian–Ediacaran) deposits in East and North-East Greenland. Memoirs, 36(1), 581-592. https://doi.org/10.1144/m36.56
• Hughes, N., Myrow, P., McKenzie, N., Harper, D., Bhargava, O., Tangri, S., …Fanning, C. (2011). Cambrian rocks and faunas of the Wachi La, Black Mountains, Bhutan. Geological Magazine, 148(03), 351-379. https://doi.org/10.1017/s0016756810000750
• of the Chernobyl NPP (Ukraine)?. Environmentalist, 31(4), 369-375. https://doi.org/10.1007/s10669-011-9347-4
• Frisk, Å., & Harper, D. (2011). Palaeoenvironmental aspects of Late Ordovician Sericoidea shell concentrations in an impact crater, Tvaren, Sweden. Lethaia: An International Journal of Palaeontology and Stratigraphy, 44(4), 383-396. https://doi.org/10.1111/j.1502-3931.2010.00250.x
• Zuykov, M., Harper, D., & Pelletier, E. (2011). Revision of the Ordovician brachiopod genus Noetlingia Hall and Clarke, 1893. Journal of Paleontology, 85(3), 595-598
• Vinther, J., Smith, M., & Harper, D. (2011). Vetulicolians from the Lower Cambrian Sirius Passet Lagerstatte, North Greenland, and the polarity of morphological characters in basal deuterostomes. Palaeontology, 54(3), 711-719. https://doi.org/10.1111/j.1475-4983.2011.01034.x
• Munnecke, A., Calner, M., & Harper, D. (2010). How does sea level correlate with sea-water chemistry? A progress report from the Ordovician and Silurian. Palaeogeography, Palaeoclimatology, Palaeoecology, 296(3-4), 213-216. https://doi.org/10.1016/j.palaeo.2010.07.009
• Rasmussen, C., Ebbestad, J., & Harper, D. (2010). Unravelling a Late Ordovician pentameride (Brachiopoda) hotspot from the Boda Limestone, Siljan district, central Sweden. GFF, 132(3-4), 133-152. https://doi.org/10.1080/11035897.2010.506008
• Azmy, K., Stouge, S., Christiansen, J., Harper, D., Knight, I., & Boyce, D. (2010). Carbon-isotope stratigraphy of the Lower Ordovician succession in Northeast Greenland: Implications for correlations with St. George Group in western Newfoundland (Canada) and beyond. Sedimentary Geology, 225(1-2), 67-81. https://doi.org/10.1016/j.sedgeo.2010.01.007
• Candela, Y., & Harper, D. (2010). Late Ordovician (Katian) brachiopods from the Southern Uplands of Scotland: biogeographic patterns on the edge of Laurentia. Earth and Environmental Science Transactions of the Royal Society of Edinburgh, 100(03), 253-274. https://doi.org/10.1017/s1755691010008121
• Munnecke, A., Calner, M., Harper, D., & Servais, T. (2010). Ordovician and Silurian sea-water chemistry, sea level, and climate: A synopsis. Palaeogeography, Palaeoclimatology, Palaeoecology, 296(3-4), 389-413. https://doi.org/10.1016/j.palaeo.2010.08.001
• Servais, T., Owen, A., Harper, D., Kroger, B., & Munnecke, A. (2010). The Great Ordovician Biodiversification Event (GOBE): The palaeoecological dimension. Palaeogeography, Palaeoclimatology, Palaeoecology, 294(3-4), 99-119. https://doi.org/10.1016/j.palaeo.2010.05.031
• Bing, H., Harper, D., Renbin, Z., & Jiayu, R. (2010). Can the Lilliput Effect be detected in the brachiopod faunas of South China following the terminal Ordovician mass extinction?. Palaeogeography, Palaeoclimatology, Palaeoecology, 285(3-4), 277-286. https://doi.org/10.1016/j.palaeo.2009.11.020
• Dahlqvist, P., Harper, D., & Wickstrom, L. (2010). Late Ordovician shelly faunas from Jamtland: palaeocommunity development along the margin of the Swedish Caledonides. Bulletin of Geosciences (On-line), 85, 505-512. https://doi.org/10.3140/bull.geosci.1175
• Harper, D., Parkes, M., & McConnell, B. (2010). Late Ordovician (Sandbian) brachiopods from the Mweelrea Formation, South Mayo, western Ireland: stratigraphic and tectonic implications. Geological Journal, 45, 445-450. https://doi.org/10.1002/gj.1210
• Jin, J., Harper, D., & Rasmussen, C. (2009). Sulcipentamerus (Pentamerida, Brachiopoda) from the Lower Silurian Washington Land Group, North Greenland. Palaeontology, 52(2), 385-399. https://doi.org/10.1111/j.1475-4983.2009.00849.x
• Rasmussen, C., Nielsen, A., & Harper, D. (2009). Ecostratigraphical interpretation of lower Middle Ordovician East Baltic sections based on brachiopods. Geological Magazine, 146(05), 717-731. https://doi.org/10.1017/s0016756809990148
• Harper, D., & Pickerill, R. (2008). Generation of brachiopod-dominated shell beds in the Miocene rocks of Carriacou, Lesser Antilles. Geological Journal, 43(5), 573-581. https://doi.org/10.1002/gj.1131
• Schmitz, B., Harper, D., Peucker-Ehrenbrink, B., Stouge, S., Alwmark, C., Cronholm, A., …Xiaofeng, W. (2008). Asteroid breakup linked to the Great Ordovician Biodiversification Event. Nature Geoscience, 1(1), 49-53. https://doi.org/10.1038/ngeo.2007.37