Staff profile

Research in the Molecular Ecology Group covers a wide range of studies and species, but our focus is on evolutionary process and genetic structure in natural populations. We use modern and ancient DNA markers to better understand the mechanisms that underlie the evolution of population structure and population dynamics in the context of ecological factors and changing environments. We use similar approaches to investigate the relationship between genetic diversity and fitness, and to better understand the role of individual behaviour in these processes. We apply this work to the development of more effective conservation strategies, and to improve our understanding of the origin and evolution of biodiversity within and among populations.

• conservation genetics
• evolutionary biology
• population genetics and molecular ecology
• Ancient DNA

• 2000: Editorial Board member: Aquatic Mammals (2005-), Open Ecology Journal (2008-), Marine Genomics (2008-), International Journal of Evolutionary Biology (2009-), Proceedings of the Royal Society, Series B (2000 - 2007) & Biology Letters (2003-2005).
• 2000: Journal Editor: Editor in Chief & Founding Editor of Conservation Genetics, Springer Publishing (2000 -). Founding Editor of Conservation Genetics Resources, Springer Publishing (2009-). Associate editor for Marine Biology Research (2006-) & Marine Mammal Science (2004 - 2007)

Authored book

• Minke Whales
  
  Hoelzel, A., & Stern, S. (2000). Minke Whales. Colin Baxter.
• Molecular Genetic Ecology
  
  Hoelzel, A., & Dover, G. (1991). Molecular Genetic Ecology. Oxford University Press.

Edited book

• Marine Mammal Biology; an Evolutionary Approach.
  
  Hoelzel, A. (Ed.). (2002). Marine Mammal Biology; an Evolutionary Approach. Blackwell.
• Molecular Epidemiology; a Practical Approach.
  
  Carrington, M., & Hoelzel, A. (Eds.). (2001). Molecular Epidemiology; a Practical Approach. Oxford University Press.
• Molecular Genetic Analysis of Populations; a Practical Approach.
  
  Hoelzel, A. (Ed.). (1998). Molecular Genetic Analysis of Populations; a Practical Approach. Oxford University Press.
• Molecular Genetic Analysis of Populations; a Practical Approach.
  
  Hoelzel, A. (Ed.). (1992). Molecular Genetic Analysis of Populations; a Practical Approach. Oxford University Press.
• Genetic Ecology of Whales and Dolphins.
  
  Hoelzel, A. (Ed.). (1991). Genetic Ecology of Whales and Dolphins. International Whaling Commission.

Journal Article

• Postglacial Recolonization of the Southern Ocean by Elephant Seals Occurred From Multiple Glacial Refugia
  
  Berg, A. A., Askew, M., Seersholm, F. V., Verry, A. J. F., Hoelzel, A. R., Welch, A., Greig, K., Walter, R., Knapp, M., Barlow, A., Paijmans, J. L. A., Waters, J. M., Bunce, M., McDonald, K., O’Connor, S., Hall, B., Koch, P. L., Baroni, C., Salvatore, M. C., … de Bruyn, M. (2025). Postglacial Recolonization of the Southern Ocean by Elephant Seals Occurred From Multiple Glacial Refugia. Global Change Biology, 31(3), Article e70101. https://doi.org/10.1111/gcb.70101
• Genomics of post-bottleneck recovery in the northern elephant seal.
  
  Hoelzel, A. R., Gkafas, G. A., Kang, H., Sarigol, F., Le Boeuf, B., Costa, D. P., Beltran, R. S., Reiter, J., Robinson, P. W., McInerney, N., Seim, I., Sun, S., Fan, G., & Li, S. (2024). Genomics of post-bottleneck recovery in the northern elephant seal. Nature Ecology & Evolution, 8, 686-694. https://doi.org/10.1038/s41559-024-02337-4
• Ancient and modern DNA track temporal and spatial population dynamics in the European fallow deer since the Eemian interglacial
  
  Baker, K. H., Gray, H. W. I., Lister, A. M., Spassov, N., Welch, A. J., Trantalidou, K., De Cupere, B., Bonillas, E., De Jong, M., Çakırlar, C., Sykes, N., & Hoelzel, A. R. (2024). Ancient and modern DNA track temporal and spatial population dynamics in the European fallow deer since the Eemian interglacial. Scientific Reports, 14, Article 3015. https://doi.org/10.1038/s41598-023-48112-6
• Moderating the neutralist–selectionist debate: exactly which propositions are we debating, and which arguments are valid?
  
  de Jong, M. J., van Oosterhout, C., Hoelzel, A. R., & Janke, A. (2024). Moderating the neutralist–selectionist debate: exactly which propositions are we debating, and which arguments are valid?. Biological Reviews, 99(1), 23-55. https://doi.org/10.1111/brv.13010
• Intraspecific phenotypic differentiation by habitat depth in deep demersal fish species
  
  Steeds, N., Zulqurnain, Z., Regnier, T., Gibb, F., Stirling, D., & Hoelzel, A. R. (2024). Intraspecific phenotypic differentiation by habitat depth in deep demersal fish species. Frontiers in Marine Science, 11, Article 1437952. https://doi.org/10.3389/fmars.2024.1437952
• Kinship study reveals stable non-kin-based associations in a medium-sized delphinid
  
  Hartman, K. L., Chen, I., van der Harst, P. A., Moura, A. E., Jahnke, M., Pilot, M., Vilela, R., & Hoelzel, A. R. (2023). Kinship study reveals stable non-kin-based associations in a medium-sized delphinid. Behavioral Ecology and Sociobiology, 77(12), Article 137. https://doi.org/10.1007/s00265-023-03411-w
• Where to now with the evolutionarily significant unit?
  
  Hoelzel, A. R. (2023). Where to now with the evolutionarily significant unit?. Trends in Ecology & Evolution, 38(12), 1134-1142. https://doi.org/10.1016/j.tree.2023.07.005
• The raw material of evolution
  
  Hoelzel, A. R., & Lynch, M. (2023). The raw material of evolution. Science, 381(6661), 942-943. https://doi.org/10.1126/science.adk0121
• Genomics reveals the role of admixture in the evolution of structure among sperm whale populations within the Mediterranean Sea
  
  Violi, B., de Jong, M. J., Frantzis, A., Alexiadou, P., Tardy, C., Ody, D., de Stephanis, R., Giménez, J., Lucifora, G., e Silva, M. A., Oliveira, C., Alves, F., Dinis, A., Tejedor, M., Fernández, A., Arregui, M., Arbelo, M., Lopez, A., Covelo, P., & Hoelzel, A. R. (2023). Genomics reveals the role of admixture in the evolution of structure among sperm whale populations within the Mediterranean Sea. Molecular Ecology, 32(11), 2715-2731. https://doi.org/10.1111/mec.16898
• Widespread southern elephant seal occupation of the Victoria land coast implies a warmer-than-present Ross Sea in the mid-to-late Holocene
  
  Hall, B. L., Koch, P. L., Baroni, C., Salvatore, M. C., Hoelzel, A. R., de Bruyn, M., & Welch, A. J. (2023). Widespread southern elephant seal occupation of the Victoria land coast implies a warmer-than-present Ross Sea in the mid-to-late Holocene. Quaternary Science Reviews, 303, Article 107991. https://doi.org/10.1016/j.quascirev.2023.107991
• Museum specimens of a landlocked pinniped reveal recent loss of genetic diversity and unexpected population connections
  
  Heino, M. T., Nyman, T., Palo, J. U., Harmoinen, J., Valtonen, M., Pilot, M., Översti, S., Salmela, E., Kunnasranta, M., Väinölä, R., Hoelzel, A. R., & Aspi, J. (2023). Museum specimens of a landlocked pinniped reveal recent loss of genetic diversity and unexpected population connections. Ecology and Evolution, 13(1), Article 9720. https://doi.org/10.1002/ece3.9720
• Strong and lasting impacts of past global warming on baleen whales and their prey
  
  Cabrera, A. A., Schall, E., Bérubé, M., Anderwald, P., Bachmann, L., Berrow, S., Best, P. B., Clapham, P. J., Cunha, H., Dalla Rosa, L., Dias, C., Findlay, K., Haug, T., Heide‐Jørgensen, M. P., Hoelzel, A., Kovacs, K. M., Landry, S., Larsen, F., Lopes, X. M., … Palsbøll, P. J. (2022). Strong and lasting impacts of past global warming on baleen whales and their prey. Global Change Biology, 28(8), 2657-2677. https://doi.org/10.1111/gcb.16085
• Evolutionary drivers of morphological differentiation among three bottlenose dolphin lineages, Tursiops spp. (Delphinidae), in the northwest Indian Ocean utilizing linear and geometric morphometric techniques
  
  Gray, H., van Waerebeek, K., Owen, J., Collins, T., Minton, G., Ponnampalam, L., Willson, A., Baldwin, R., & Hoelzel, A. R. (2022). Evolutionary drivers of morphological differentiation among three bottlenose dolphin lineages, Tursiops spp. (Delphinidae), in the northwest Indian Ocean utilizing linear and geometric morphometric techniques. Biological Journal of the Linnean Society, 135(3), 610-629. https://doi.org/10.1093/biolinnean/blab133
• Population structure associated with bioregion and seasonal prey distribution for Indo‐Pacific bottlenose dolphins (Tursiops aduncus) in South Africa
  
  Vargas‐Fonseca, O. A., Yates, P., Kirkman, S. P., Pistorius, P. A., Moore, D. M., Natoli, A., Cockcroft, V., & Hoelzel, A. R. (2021). Population structure associated with bioregion and seasonal prey distribution for Indo‐Pacific bottlenose dolphins (Tursiops aduncus) in South Africa. Molecular Ecology, 30(19), 4642-4659. https://doi.org/10.1111/mec.16086
• Comparative genomics provides insights into the aquatic adaptations of mammals
  
  Yuan, Y., Zhang, Y., Zhang, P., Liu, C., Wang, J., Gao, H., Hoelzel, A. R., Seim, I., Lv, M., Lin, M., Dong, L., Gao, H., Yang, Z., Caruso, F., Lin, W., da Fonseca, R. R., Wang, D., Wang, X., Rasmussen, M. H., … Li, S. (2021). Comparative genomics provides insights into the aquatic adaptations of mammals. Proceedings of the National Academy of Sciences, 118(37), Article e2106080118. https://doi.org/10.1073/pnas.2106080118
• Conservation genetics of the European fallow deer: a reply to Marchesini et al
  
  Baker, K. H., Gray, H. W., Pekşen, Çiğdem A., & Hoelzel, A. R. (2021). Conservation genetics of the European fallow deer: a reply to Marchesini et al. Mammalian Biology, 101(3), 313-319. https://doi.org/10.1007/s42991-021-00121-6
• Marine mammal conservation: over the horizon
  
  Nelms, S., Alfaro-Shigueto, J., Arnould, J., Avila, I., Bengtson Nash, S., Campbell, E., Carter, M., Collins, T., Currey, R., Domit, C., Franco-Trecu, V., Fuentes, M., Gilman, E., Harcourt, R., Hines, E., Hoelzel, A., Hooker, S., Johnston, D., Kelkar, N., … Godley, B. (2021). Marine mammal conservation: over the horizon. Endangered Species Research, 44, 291-325. https://doi.org/10.3354/esr01115
• Comparative biogeography and the evolution of population structure for bottlenose and common dolphins in the Indian Ocean
  
  Gray, H. W., Chen, I., Moura, A. E., Natoli, A., Nishida, S., Tanabe, S., Minton, G., Ponnampalam, L. S., Kiani, M. S., Culloch, R., Gore, M., Särnblad, A., Amir, O., Berggren, P., Collins, T., Willson, A. J., Baldwin, R., & Hoelzel, A. R. (2021). Comparative biogeography and the evolution of population structure for bottlenose and common dolphins in the Indian Ocean. Journal of Biogeography, 48(7). https://doi.org/10.1111/jbi.14102
• Genomic signatures of divergent selection are associated with social behaviour for spinner dolphin ecotypes
  
  Andrews, K. R., Epstein, B., Leslie, M. S., Fiedler, P., Morin, P. A., & Hoelzel, A. R. (2021). Genomic signatures of divergent selection are associated with social behaviour for spinner dolphin ecotypes. Molecular Ecology, 30(9). https://doi.org/10.1111/mec.15865
• Demography and adaptation promoting evolutionary transitions in a mammalian genus that diversified during the Pleistocene
  
  De Jong, M., Li, Z., Qin, Y., Quemere, E., Baker, K., Wang, W., & Hoelzel, A. (2020). Demography and adaptation promoting evolutionary transitions in a mammalian genus that diversified during the Pleistocene. Molecular Ecology, 29(15), 2777-2792. https://doi.org/10.1111/mec.15450
• Population genetic diversity and historical dynamics of Fraser’s dolphins Lagenodelphis hosei
  
  Chen, I., Nishida, S., Chou, L., Isobe, T., Mignucci-Giannoni, A., & Hoelzel, A. (2020). Population genetic diversity and historical dynamics of Fraser’s dolphins Lagenodelphis hosei. Marine Ecology Progress Series, 643, 183-195. https://doi.org/10.3354/meps13268
• Sex-specific impact of inbreeding on pathogen load in the striped dolphin
  
  Gkafas, G. A., de Jong, M., Exadactylos, A., Raga, J. A., Aznar, F. J., & Hoelzel, A. R. (2020). Sex-specific impact of inbreeding on pathogen load in the striped dolphin. Proceedings of the Royal Society B: Biological Sciences, 287(1922), Article 20200195. https://doi.org/10.1098/rspb.2020.0195
• Genomic data suggest environmental drivers of fish population structure in the deep sea: A case study for the orange roughy (Hoplostethus atlanticus)
  
  Gonçalves da Silva, A., Barendse, W., Kijas, J., England, P. R., & Hoelzel, A. R. (2020). Genomic data suggest environmental drivers of fish population structure in the deep sea: A case study for the orange roughy (Hoplostethus atlanticus). Journal of Applied Ecology, 57(2), 296-306. https://doi.org/10.1111/1365-2664.13534
• A global cline in a colour polymorphism suggests a limited contribution of gene flow towards the recovery of a heavily exploited marine mammal
  
  Hoffman, J., Bauer, E., Paijmans, A., Humble, E., Beckmann, L., Kubetschek, C., Christaller, F., Kröcker, N., Fuchs, B., Moreras, A., Shihlomule, Y., Bester, M., Cleary, A., De Bruyn, P., Forcada, J., Goebel, M., Goldsworthy, S., Guinet, C., Hoelzel, A., … Lowther, A. (2018). A global cline in a colour polymorphism suggests a limited contribution of gene flow towards the recovery of a heavily exploited marine mammal. Royal Society Open Science, 5(10), Article 181227. https://doi.org/10.1098/rsos.181227
• Hybridization in bottlenose dolphins—A case study of Tursiops aduncus × T. truncatus hybrids and successful backcross hybridization events
  
  Gridley, T., Elwen, S., Harris, G., Moore, D., Hoelzel, A., & Lampen, F. (2018). Hybridization in bottlenose dolphins—A case study of Tursiops aduncus × T. truncatus hybrids and successful backcross hybridization events. PLoS ONE, 13(9), Article e0201722. https://doi.org/10.1371/journal.pone.0201722
• Eco-Evolutionary Processes Generating Diversity Among Bottlenose Dolphin, Tursiops truncatus, Populations off Baja California, Mexico
  
  Segura-García, I., Rojo-Arreola, L., Rocha-Olivares, A., Heckel, G., Gallo-Reynoso, J. P., & Hoelzel, R. (2018). Eco-Evolutionary Processes Generating Diversity Among Bottlenose Dolphin, Tursiops truncatus, Populations off Baja California, Mexico. Evolutionary Biology, 45(2), 223-236. https://doi.org/10.1007/s11692-018-9445-z
• Cryptic Lineage Differentiation Among Indo-Pacific Bottlenose Dolphins (Tursiops aduncus) in the Northwest Indian Ocean
  
  Gray, H., Nishida, S., Welch, A., Moura, A., Tanabe, S., Kiani, M., Culloch, R., Möller, L., Natoli, A., Ponnampalam, L., Minton, G., Gore, M., Collins, T., Willson, A., Baldwin, R., & Hoelzel, A. (2018). Cryptic Lineage Differentiation Among Indo-Pacific Bottlenose Dolphins (Tursiops aduncus) in the Northwest Indian Ocean. Molecular Phylogenetics and Evolution, 122, 1-14. https://doi.org/10.1016/j.ympev.2017.12.027
• Genomics of habitat choice and adaptive evolution in a deep-sea fish
  
  Gaither, M. R., Gkafas, G. A., de Jong, M., Sarigol, F., Neat, F., Regnier, T., Moore, D., Grӧcke D. R., Hall, N., Liu, X., Kenny, J., Lucaci, A., Hughes, M., Haldenby, S., & Hoelzel, A. R. (2018). Genomics of habitat choice and adaptive evolution in a deep-sea fish. Nature Ecology and Evolution, 2(4), 680-687. https://doi.org/10.1038/s41559-018-0482-x
• Biogeography and temporal progression during the evolution of striped dolphin population structure in European waters
  
  Gkafas, G., Exadactylos, A., Rogan, E., Raga, J., Reid, R., & Hoelzel, A. (2017). Biogeography and temporal progression during the evolution of striped dolphin population structure in European waters. Journal of Biogeography, 44(12), 2681-2691. https://doi.org/10.1111/jbi.13079
• Genetic diversity of bottlenose dolphin (Tursiops sp.) populations in the western North Pacific and the conservation implications
  
  Chen, I., Nishida, S., Yang, W.-C., Isobe, T., Tajima, Y., & Hoelzel, A. R. (2017). Genetic diversity of bottlenose dolphin (Tursiops sp.) populations in the western North Pacific and the conservation implications. Marine Biology, 164(10), Article 202. https://doi.org/10.1007/s00227-017-3232-8
• Ancient female philopatry, asymmetric male gene flow, and synchronous population expansion support the influence of climatic oscillations on the evolution of South American sea lion (Otaria flavescens)
  
  Oliveira, L. R. de, Gehara, M. C., Fraga, L. D., Lopes, F., Túnez, J. I., Cassini, M. H., Majluf, P., Cárdenas-Alayza, S., Pavés, H. J., Crespo, E. A., García, N., Loizaga de Castro, R., Hoelzel, A. R., Sepúlveda, M., Olavarría, C., Valiati, V. H., Quiñones, R., Pérez-Alvarez, M. J., Ott, P. H., & Bonatto, S. L. (2017). Ancient female philopatry, asymmetric male gene flow, and synchronous population expansion support the influence of climatic oscillations on the evolution of South American sea lion (Otaria flavescens). PLoS ONE, 12(6), Article e0179442. https://doi.org/10.1371/journal.pone.0179442
• Strong population structure in a species manipulated by humans since the Neolithic: The European fallow deer (Dama dama dama)
  
  Baker, K., Gray, H., Ramovs, V., Mertzanidou, D., Akın Pekşen, Ç, Bilgin, C., Sykes, N., & Hoelzel, A. (2017). Strong population structure in a species manipulated by humans since the Neolithic: The European fallow deer (Dama dama dama). Heredity, 119, 16-26. https://doi.org/10.1038/hdy.2017.11
• Depth as a driver of evolution in the deep sea: Insights from grenadiers (Gadiformes: Macrouridae) of the genus Coryphaenoides
  
  Gaither, M., Violia, B., Gray, H., Neat, F., Drazen, J., Grubbs, D., Roa-Varón, A., Sutton, T., & Hoelzel, A. R. (2016). Depth as a driver of evolution in the deep sea: Insights from grenadiers (Gadiformes: Macrouridae) of the genus Coryphaenoides. Molecular Phylogenetics and Evolution, 104, 73-82. https://doi.org/10.1016/j.ympev.2016.07.027
• Population differentiation in the context of Holocene climate change for a migratory marine species, the southern elephant seal
  
  Corrigan, L., Fabiani, A., Chauke, L., McMahon, C., de Bruyn, M., Bester, M., Bastos, A., Campagna, C., Muelbert, M., & Hoelzel, A. (2016). Population differentiation in the context of Holocene climate change for a migratory marine species, the southern elephant seal. Journal of Evolutionary Biology, 29(9), 1667-1679. https://doi.org/10.1111/jeb.12870
• Phylogenomics of the killer whale indicates ecotype divergence in sympatry
  
  Moura, A., Kenny, J., Chaudhuri, R., Hughes, M., Reisinger, R., de Bruyn, P., Dahlheim, M., Hall, N., & Hoelzel, A. (2015). Phylogenomics of the killer whale indicates ecotype divergence in sympatry. Heredity, 114(1), 48-55. https://doi.org/10.1038/hdy.2014.67
• Delphinid systematics and biogeography with a focus on the current genus Lagenorhynchus: Multiple pathways for antitropical and trans-oceanic radiation
  
  Banguera-Hinestroza, E., Hayano, A., Crespo, E., & Hoelzel, A. (2014). Delphinid systematics and biogeography with a focus on the current genus Lagenorhynchus: Multiple pathways for antitropical and trans-oceanic radiation. Molecular Phylogenetics and Evolution, 80, 217-230. https://doi.org/10.1016/j.ympev.2014.08.005
• Population genomics of the killer whale indicates ecotype evolution in sympatry involving both selection and drift
  
  Moura, A., Kenny, J., Chaudhuri, R., Hughes, M., Welch, A., Reisinger, R., de Bruyn, P., Dahlheim, M., Hall, N., & Hoelzel, A. (2014). Population genomics of the killer whale indicates ecotype evolution in sympatry involving both selection and drift. Molecular Ecology, 23(21), 5179-5192. https://doi.org/10.1111/mec.12929
• Influence of Holocene environmental change and anthropogenic impact on the diversity and distribution of roe deer
  
  Baker, K., & Hoelzel, A. (2014). Influence of Holocene environmental change and anthropogenic impact on the diversity and distribution of roe deer. Heredity, 112(6), 607-615. https://doi.org/10.1038/hdy.2013.142
• Killer Whale Nuclear Genome and mtDNA Reveal Widespread Population Bottleneck During the Last Glacial Maximum
  
  Moura, A., Janse van Rensburg, C., Pilot, M., Tehrani, A., Best, P., Thornton, M., Plön, S., de Bruyn, P., Worley, K., Gibbs, R., Dahlheim, M., & Hoelzel, A. (2014). Killer Whale Nuclear Genome and mtDNA Reveal Widespread Population Bottleneck During the Last Glacial Maximum. Molecular Biology and Evolution, 31(5), 1121-1131. https://doi.org/10.1093/molbev/msu058
• Rapid increase in southern elephant seal genetic diversity after a founder event
  
  De Bruyn, M., Pinsky, M., Hall, B., Koch, P., Baroni, C., & Hoelzel, A. (2014). Rapid increase in southern elephant seal genetic diversity after a founder event. Proceedings of the Royal Society B: Biological Sciences, 281(1779), Article 20133078. https://doi.org/10.1098/rspb.2013.3078
• Recent diversification of a marine genus (Tursiops spp.) tracks habitat preference and environmental change
  
  Moura, A., Nielsen, S., Vilstrup, J., Moreno-Mayar, J., Gilbert, M., Gray, H., Natoli, A., Möller, L., & Hoelzel, A. (2013). Recent diversification of a marine genus (Tursiops spp.) tracks habitat preference and environmental change. Systematic Biology, 62(6), 865-877. https://doi.org/10.1093/sysbio/syt051
• Does Presence of a Mid-Ocean Ridge Enhance Biomass and Biodiversity?
  
  Priede, I. G., Bergstad, O. A., Miller, P. I., Vecchione, M., Gebruk, A., Falkenhaug, T., Billett, D. S., Craig, J., Dale, A. C., Shields, M. A., Tilstone, G. H., Sutton, T. T., Gooday, A. J., Inall, M. E., Jones, D. O., Martinez-Vicente, V., Menezes, G. M., Niedzielski, T., Sigurðsson, Þorsteinn, … Lorance, P. (2013). Does Presence of a Mid-Ocean Ridge Enhance Biomass and Biodiversity?. PLoS ONE, 8(5), Article e61550. https://doi.org/10.1371/journal.pone.0061550
• Phenotypic and genetic divergence among harbour porpoise populations associated with habitat regions in the North Sea and adjacent seas.
  
  De Luna, C., Goodman, S., Thatcher, O., Jepson, P., Andersen, L., Tolley, K., & Hoelzel, A. (2012). Phenotypic and genetic divergence among harbour porpoise populations associated with habitat regions in the North Sea and adjacent seas. Journal of Evolutionary Biology, 25(4), 674-681. https://doi.org/10.1111/j.1420-9101.2012.02461.x
• Spatial Scale and Environmental Determinants in Minke Whale Habitat Use and Foraging.
  
  Anderwald, P., Evans, P., Dyer, R., Dale, A., Wright, P., & Hoelzel, A. (2012). Spatial Scale and Environmental Determinants in Minke Whale Habitat Use and Foraging. Marine Ecology Progress Series, 450, 259-274. https://doi.org/10.3354/meps09573
• Kinship of long-term associates in the highly social sperm whale.
  
  Ortega-Ortiz, J., Engelhaupt, D., Winsor, M., Mate, B., & Hoelzel, A. (2012). Kinship of long-term associates in the highly social sperm whale. Molecular Ecology, 21(3), 732-744. https://doi.org/10.1111/j.1365-294x.2011.05274.x
• Environmental factors associated with genetic and phenotypic divergence among sympatric populations of Arctic charr (Salvelinus alpinus)
  
  Corrigan, L., Lucas, M., Winfield, I., & Hoelzel, A. (2011). Environmental factors associated with genetic and phenotypic divergence among sympatric populations of Arctic charr (Salvelinus alpinus). Journal of Evolutionary Biology, 24(9), 1906-1917. https://doi.org/10.1111/j.1420-9101.2011.02327.x
• Dietary plasticity in Arctic charr (Salvelinus alpinus) in response to long-term environmental change
  
  Corrigan, L., Winfield, I., Hoelzel, A., & Lucas, M. (2011). Dietary plasticity in Arctic charr (Salvelinus alpinus) in response to long-term environmental change. Ecology of Freshwater Fish, 20(1), 5-13. https://doi.org/10.1111/j.1600-0633.2010.00446.x
• The impact of population bottlenecks on fluctuating asymmetry and morphological variance in two separate populations of reindeer on the island of South Georgia
  
  Lovatt, F., & Hoelzel, A. (2011). The impact of population bottlenecks on fluctuating asymmetry and morphological variance in two separate populations of reindeer on the island of South Georgia. Biological Journal of the Linnean Society, 102(4), 798-811. https://doi.org/10.1111/j.1095-8312.2011.01600.x
• Comparative assessment of population genetics and demographic history of two congeneric deep sea fish species living at different depths.
  
  White, T., Fotherby, H., & Hoelzel, A. (2011). Comparative assessment of population genetics and demographic history of two congeneric deep sea fish species living at different depths. Marine Ecology Progress Series, 434, 155-164. https://doi.org/10.3354/meps09207
• Faunal histories from Holocene ancient DNA.
  
  deBruyn, M., Hoelzel, A., Carvalho, G., & Hofreiter, M. (2011). Faunal histories from Holocene ancient DNA. Trends in Ecology and Evolution, 26(8), 405-413. https://doi.org/10.1016/j.tree.2011.03.021
• Genetic panmixia and demographic dependence across the North Atlantic in the deep-sea fish, blue hake (Antimora rostrata).
  
  White, T., Fotherby, H., Stephens, P., & Hoelzel, A. (2011). Genetic panmixia and demographic dependence across the North Atlantic in the deep-sea fish, blue hake (Antimora rostrata). Heredity, 106(4), 690-699. https://doi.org/10.1038/hdy.2010.108
• Isolation and characterization of ten nuclear microsatellite DNA loci for the Portuguese dogfish Centroscymnus coelolepis.
  
  Verissimo, A., Moura, T., McDowell, J., Graves, J., Gordo, L., & Hoelzel, A. (2011). Isolation and characterization of ten nuclear microsatellite DNA loci for the Portuguese dogfish Centroscymnus coelolepis. Conservation Genetics Resources, 3(2). https://doi.org/10.1007/s12686-010-9347-6
• Seabird feeding strategies and their role in predicting association patterns with minke whales.
  
  Anderwald, P., Evans, P., Lorenz, G., & Hoelzel, A. (2011). Seabird feeding strategies and their role in predicting association patterns with minke whales. Marine Ecology Progress Series, 424, 219-227. https://doi.org/10.3354/meps08947
• Possible cryptic stock structure for minke whales in the North Atlantic; Implications for conservation and management.
  
  Anderwald, P., Danielsdottir, A., Haug, T., Laren, F., Lesage, V., Reid, R., Vikingsson, G., & Hoelzel, A. (2011). Possible cryptic stock structure for minke whales in the North Atlantic; Implications for conservation and management. Biological Conservation, 144(10), 2479-2489. https://doi.org/10.1016/j.biocon.2011.07.002
• New evidence for the establishment and management of the European fallow deer (Dama dama dama) in Roman Britain.
  
  Sykes, N., Baker, K., Carden, R., Higham, T., Hoelzel, A., & Stevens, R. (2011). New evidence for the establishment and management of the European fallow deer (Dama dama dama) in Roman Britain. Journal of Archaeological Science, Reports., 38(1), 156-165. https://doi.org/10.1016/j.jas.2010.08.024
• Insights into genetic diversity, parentage and group composition of Atlantic white-sided dolphins (Lagenorhynchus acutus) in the west of Ireland based on nuclear and mitochondrial genetic markers.
  
  Mirimin, L., Banguera-Hinestroza, E., Hoelzel, A., R, C., !&, T., & gan, E. (2011). Insights into genetic diversity, parentage and group composition of Atlantic white-sided dolphins (Lagenorhynchus acutus) in the west of Ireland based on nuclear and mitochondrial genetic markers. Journal of Heredity, 102(1), 79-87. https://doi.org/10.1093/jhered/esq106
• Local selection and population structure in a deep-sea fish, the roundnose grenadier (Coryphaenoides rupestris).
  
  White, T., Stamford, J., & Hoelzel, A. (2010). Local selection and population structure in a deep-sea fish, the roundnose grenadier (Coryphaenoides rupestris). Molecular Ecology, 19(2), 216-226. https://doi.org/10.1111/j.1365-294x.2009.04446.x
• Southernmost distribution of common bottlenose dolphins (Tursiops truncatus) in the eastern South Pacific.
  
  Olavarria, C., Acevedo, J., Vester, H., Zamorano-Abramson, J., Viddi, F., Gibbins, J., Newcombe, E., Capella, J., Hoelzel, A., Flores, F., Hucke-Gaete, R., & Torres-Florenz, J. (2010). Southernmost distribution of common bottlenose dolphins (Tursiops truncatus) in the eastern South Pacific. Aquatic Mammals., 36(3). https://doi.org/10.1578/am.36.3.2010.288
• Nineteen new microsatellite loci for the blue hake (Antimoura rostrata).
  
  White, T., Fotherby, H., & Hoelzel, A. (2010). Nineteen new microsatellite loci for the blue hake (Antimoura rostrata). Conservation Genetics Resources, 2(S1). https://doi.org/10.1007/s12686-009-9164-y
• Social cohesion among kin, gene flow without dispersal and the evolution of population genetic structure in the killer whale (Orcinus orca).
  
  Pilot, M., Dahlheim, M., & Hoelzel, A. (2010). Social cohesion among kin, gene flow without dispersal and the evolution of population genetic structure in the killer whale (Orcinus orca). Journal of Evolutionary Biology, 23(1), 20-31. https://doi.org/10.1111/j.1420-9101.2009.01887.x
• The influence of glacial epochs and habitat dependence on the diversity and phylogeography of a coastal dolphin species: Lagenorhynchus albirostris.
  
  Banguera-Hinestroza, E., Bjørge, A., Reid, R., Jepson, P., & Hoelzel, A. (2010). The influence of glacial epochs and habitat dependence on the diversity and phylogeography of a coastal dolphin species: Lagenorhynchus albirostris. Conservation Genetics, 11(5), 1823-1836. https://doi.org/10.1007/s10592-010-0075-y
• Looking backwards to look forwards : conservation genetics in a changing world.
  
  Hoelzel, A. (2010). Looking backwards to look forwards : conservation genetics in a changing world. Conservation Genetics, 11(2), 655-660. https://doi.org/10.1007/s10592-010-0045-4
• Development of 10 microsatellite loci in the ling (Molva molva)
  
  Ring, A., Knutsen, H., Fiani, D., Hoelzel, A., & Andre, C. (2009). Development of 10 microsatellite loci in the ling (Molva molva). Molecular Ecology Resources, 9(5), 1401-1403.
• Bathymetric barriers promoting genetic structure in the deepwater demersal fish tusk (Brosme brosme)
  
  Knutsen, H., Jorde, P., Sannaes, H., Hoelzel, A., Bergstad, O., Stefanni, S., Johansen, T., & Stenseth, N. (2009). Bathymetric barriers promoting genetic structure in the deepwater demersal fish tusk (Brosme brosme). Molecular Ecology, 18(15), 3151-3162.
• Population genetics of an invasive riparian species, Impatiens glandulifera
  
  Walker, N., Hulme, P., & Hoelzel, A. (2009). Population genetics of an invasive riparian species, Impatiens glandulifera. Plant Ecology, 203(2), 243-252.
• Rapid Response of a Marine Mammal Species to Holocene Climate and Habitat Change
  
  de Bruyn, M., Hall, B., Chauke, L., Baroni, C., Koch, P., & Hoelzel, A. (2009). Rapid Response of a Marine Mammal Species to Holocene Climate and Habitat Change. PLoS Genetics, 5(7), Article e1000554. https://doi.org/10.1371/journal.pgen.1000554
• Unexpected panmixia in a long-lived, deep-sea fish with well-defined spawning habitat and relatively low fecundity
  
  White, T., Stefanni, S., Stamford, J., & Hoelzel, A. (2009). Unexpected panmixia in a long-lived, deep-sea fish with well-defined spawning habitat and relatively low fecundity. Molecular Ecology, 18(12), 2563-2573. https://doi.org/10.1111/j.1365-294x.2009.04218.x
• Nine new microsatellite loci for the orange roughy (Hoplostethus atlanticus)
  
  White, T., Stamford, J., & Hoelzel, A. (2009). Nine new microsatellite loci for the orange roughy (Hoplostethus atlanticus). Conservation Genetics, 10(3), 601-603.
• Balancing and Directional Selection at Exon-2 of the MHC DQB1 Locus among Populations of Odontocete Cetaceans
  
  Vassilakos, D., Natoli, A., Dahlheim, M., & Hoelzel, A. (2009). Balancing and Directional Selection at Exon-2 of the MHC DQB1 Locus among Populations of Odontocete Cetaceans. Molecular Biology and Evolution, 26(3), 681-689. https://doi.org/10.1093/molbev/msn296
• Female philopatry in coastal basins and male dispersion across the North Atlantic in a highly mobile marine species, the sperm whale (Physeter macrocephalus).
  
  Engelhaupt, D., Hoelzel, A., Nicholson, C., Frantzis, A., Mesnick, S., Gero, S., Whitehead, H., Rendell, L., Miller, P., De Stefanis, R., Canadas, A., Airoldi, S., & Mignucci-Giannoni, A. (2009). Female philopatry in coastal basins and male dispersion across the North Atlantic in a highly mobile marine species, the sperm whale (Physeter macrocephalus). Molecular Ecology, 18(20), 4193-4205. https://doi.org/10.1111/j.1365-294x.2009.04355.x
• Conservation genetics of the short-beaked common dolphin (Delphinus delphis) in the Mediterranean Sea and in the eastern North Atlantic Ocean
  
  Natoli, A., Canadas, A., Vaquero, C., Politi, E., Fernandez-Navarro, P., & Hoelzel, A. (2008). Conservation genetics of the short-beaked common dolphin (Delphinus delphis) in the Mediterranean Sea and in the eastern North Atlantic Ocean. Conservation Genetics, 9(6), 1479-1487.
• Isolation and characterization of microsatellite loci in the deep-sea marine fish, the roundnose grenadier (Coryphaenoides rupestris)
  
  Knutsen, H., Le Goff-Vitry, M., Fiani, D., & Hoelzel, A. (2008). Isolation and characterization of microsatellite loci in the deep-sea marine fish, the roundnose grenadier (Coryphaenoides rupestris). Molecular Ecology Resources, 8(5), 993-995.
• Population structure of bottlenose dolphins (Tursiops aduncus) impacted by bycatch along the east coast of South Africa
  
  Natoli, A., Peddemors, V., & Hoelzel, A. (2008). Population structure of bottlenose dolphins (Tursiops aduncus) impacted by bycatch along the east coast of South Africa. Conservation Genetics, 9(3), 627-636.
• Temporal and contextual patterns of killer whale (Orcinus orca) call type production
  
  Foote, A., Osborne, R., & Hoelzel, A. (2008). Temporal and contextual patterns of killer whale (Orcinus orca) call type production. Ethology, 114(6), 599-606.
• Ancient human mtDNA genotypes from England reveal lost variation over the last millennium
  
  Topf, A., Gilbert, M., Fleischer, R., & Hoelzel, A. (2007). Ancient human mtDNA genotypes from England reveal lost variation over the last millennium. Biology Letters, 3(5), 550-553.
• Isolation and characterization of microsatellite loci in a marine fish species, the tusk (Brosme brosme)
  
  Knutsen, H., Fiani, D., Sannae, H., & Hoelzel, A. (2007). Isolation and characterization of microsatellite loci in a marine fish species, the tusk (Brosme brosme). Molecular Ecology Notes., 7(5), 851-853.
• Genetic isolation of a now extinct population of bottlenose dolphins (Tursiops truncatus)
  
  Nichols, C., Herman, J., Gaggiotti, O., Dobney, K., Parsons, K., & Hoelzel, A. (2007). Genetic isolation of a now extinct population of bottlenose dolphins (Tursiops truncatus). Proceedings of the Royal Society B: Biological Sciences, 274(1618), 1611-1616.
• Social kin associations and genetic structuring of striped dolphin populations (Stenella coeruleoalba) in the Mediterranean Sea
  
  Gaspari, S., Azzellino, A., Airoldi, S., & Hoelzel, A. (2007). Social kin associations and genetic structuring of striped dolphin populations (Stenella coeruleoalba) in the Mediterranean Sea. Molecular Ecology, 16(14), 2922-2933.
• Evolution of population structure in a highly social top predator, the killer whale
  
  Hoelzel, A., Hey, J., Dahlheim, M., Nicholson, C., Burkanov, V., & Black, N. (2007). Evolution of population structure in a highly social top predator, the killer whale. Molecular Biology and Evolution, 24(6), 1407-1415. https://doi.org/10.1093/molbev/msm063
• Risso's dolphins (Grampus griseus) in UK waters are differentiated from a population in the Mediterranean Sea and genetically less diverse
  
  Gaspari, S., Airoldi, S., & Hoelzel, A. (2007). Risso’s dolphins (Grampus griseus) in UK waters are differentiated from a population in the Mediterranean Sea and genetically less diverse. Conservation Genetics, 8(3), 727-732.
• Population genetic structure in the North Atlantic Greenland halibut (Reinhardtius hippoglossoides): influenced by oceanic current systems?
  
  Knutsen, H., Jorde, P., Albert, O., Hoelzel, A., & Stenseth, N. (2007). Population genetic structure in the North Atlantic Greenland halibut (Reinhardtius hippoglossoides): influenced by oceanic current systems?. Canadian Journal of Fisheries and Aquatic Sciences, 64(6), 857-866.
• Genetic tracking of basking shark products in international trade
  
  Magnussen, J., Pikitch, E., Clarke, S., Nicholson, C., Hoelzel, A., & Shivji, M. (2007). Genetic tracking of basking shark products in international trade. Animal Conservation, 10(2), 199-207.
• Killer whales are capable of vocal learning
  
  Foote, A., Griffin, R., Howitt, D., Larsson, L., Miller, P., & Hoelzel, A. (2006). Killer whales are capable of vocal learning. Biology Letters, 2(4), 509-512.
• Low worldwide genetic diversity in the basking shark (Cetorhinus maximus)
  
  Hoelzel, A., Shivji, M., Magnussen, J., & Francis, M. (2006). Low worldwide genetic diversity in the basking shark (Cetorhinus maximus). Biology Letters, 2(4), 639-642.
• Sex-specific foraging strategies and resource partitioning in the southern elephant seal (Mirounga leonina)
  
  Lewis, R., O’Connell, T., Lewis, M., Carnpagna, C., & Hoelzel, A. (2006). Sex-specific foraging strategies and resource partitioning in the southern elephant seal (Mirounga leonina). Proceedings of the Royal Society B: Biological Sciences, 273(1603), 2901-2907.
• Holocene elephant seal distribution implies warmer-than-present climate in the Ross Sea
  
  Hall, B., Hoelzel, A., Baroni, C., Denton, G., Le Boeuf, B., Overturf, B., & Topf, A. (2006). Holocene elephant seal distribution implies warmer-than-present climate in the Ross Sea. Proceedings of the National Academy of Sciences, 103(27), 10213-10217.
• Phylogeography and alpha taxonomy of the common dolphin (Delphinus sp.)
  
  Natoli, A., Canadas, A., Peddemors, V., Aguilar, A., Vaquero, C., Fernandez-Piqueras, P., & Hoelzel, A. (2006). Phylogeography and alpha taxonomy of the common dolphin (Delphinus sp.). Journal of Evolutionary Biology, 19(3), 943-954.
• Tracing the phylogeography of human populations in Britain based on 4th-11th century mtDNA genotypes
  
  Topf, A., Gilbert, M., Dumbacher, J., & Hoelzel, A. (2006). Tracing the phylogeography of human populations in Britain based on 4th-11th century mtDNA genotypes. Molecular Biology and Evolution, 23(1), 152-161.
• Ancient genomes
  
  Hoelzel, A. (2005). Ancient genomes. Genome Biology, 6(12), Article 239. https://doi.org/10.1186/gb-2005-6-12-239
• A Romani mitochondrial haplotype in England 500 years before their recorded arrival in Britain
  
  Topf, A., & Hoelzel, A. (2005). A Romani mitochondrial haplotype in England 500 years before their recorded arrival in Britain. Biology Letters, 1(3), 280-282.
• Habitat structure and the dispersal of male and female bottlenose dolphins (Tursiops truncatus)
  
  Natoli, A., Birkun, A., Aguilar, A., Lopez, A., & Hoelzel, A. (2005). Habitat structure and the dispersal of male and female bottlenose dolphins (Tursiops truncatus). Proceedings of the Royal Society B: Biological Sciences, 272(1569), 1217-1226.
• Extreme polygyny among southern elephant seals on Sea Lion Island, Falkland Islands
  
  Fabiani, A., Galimberti, F., Sanvito, S., & Hoelzel, A. (2004). Extreme polygyny among southern elephant seals on Sea Lion Island, Falkland Islands. Behavioral Ecology, 15(6), 961-969.
• Whale-call response to masking boat noise
  
  Foote, A., Osborne, R., & Hoelzel, A. (2004). Whale-call response to masking boat noise. Nature, 428(6986), 910-910. https://doi.org/10.1038/428910a
• Population structure and speciation in the genus Tursiops based on microsatellite and mitochondrial DNA analyses
  
  Natoli, A., Peddemors, V., & Hoelzel, A. (2004). Population structure and speciation in the genus Tursiops based on microsatellite and mitochondrial DNA analyses. Journal of Evolutionary Biology, 17(2), 363-375.
• Population genetics of an invasive species, Heracleum mantegazzianum: implications for the role of life history, demographics and independent introductions
  
  Walker, N., Hulme, P., & Hoelzel, A. (2003). Population genetics of an invasive species, Heracleum mantegazzianum: implications for the role of life history, demographics and independent introductions. Molecular Ecology, 12(7), 1747-1756.
• A phylogeny of Chinese species in the genus Phrynocephalus (Agamidae) inferred from mitochondrial DNA sequences
  
  Pang, J., Wang, Y., Zhong, Y., Hoelzel, A., Papenfuss, T., Zeng, X., Ananjeva, N., & Zhang, Y. (2003). A phylogeny of Chinese species in the genus Phrynocephalus (Agamidae) inferred from mitochondrial DNA sequences. Molecular Phylogenetics and Evolution, 27(3), 398-409.
• Long-range paternal gene flow in the southern elephant seal
  
  Fabiani, A., Hoelzel, A., Galimberti, F., & Muelbert, M. (2003). Long-range paternal gene flow in the southern elephant seal. Science, 299(5607), 676-676.
• Lack of mtDNA control region variation in Hainan Eld's deer: Consequence of a recent population bottleneck?
  
  Pang, J., Hoelzel, A., Song, Y., Zeng, Z., & Zhang, Y. (2003). Lack of mtDNA control region variation in Hainan Eld’s deer: Consequence of a recent population bottleneck?. Conservation Genetics, 4(1), 109-112.
• Low world-wide genetic diversity in the killer whale (Orcinus orca); Implications for demographic history
  
  Hoelzel, A., Natoli, A., Dahlheim, M., Olavarria, C., Baird, R., & Black, N. (2002). Low world-wide genetic diversity in the killer whale (Orcinus orca); Implications for demographic history. Proceedings of the Royal Society B: Biological Sciences, 269(1499), 1467-1475. https://doi.org/10.1098/rspb.2002.2033
• Direct Evidence for the Impact of a Population Bottleneck on Symmetry and Genetic Diversity in the Northern Elephant Seal.
  
  Hoelzel, A., Fleischer, R., Campagna, C., Le Beouf, B., & Alvord, G. (2002). Direct Evidence for the Impact of a Population Bottleneck on Symmetry and Genetic Diversity in the Northern Elephant Seal. Journal of Evolutionary Biology, 15, 567-575.
• Phylogenetic relationships within the eared seals (Otariidae : Carnivora): Implications for the historical biogeography of the family
  
  Wynen, L., Goldsworthy, S., Insley, S., Adams, M., Bickham, J., Francis, J., Gallo, J., Hoelzel, A., Majluf, P., White, R., & Slade, R. (2001). Phylogenetic relationships within the eared seals (Otariidae : Carnivora): Implications for the historical biogeography of the family. Molecular Phylogenetics and Evolution, 21(2), 270-284.
• Characterization of major histocompatibility complex DRB diversity in the endemic South African antelope Damaliscus pygargus: a comparison in two subspecies with different demographic histories
  
  Van der Walt, J., Nel, L., & Hoelzel, A. (2001). Characterization of major histocompatibility complex DRB diversity in the endemic South African antelope Damaliscus pygargus: a comparison in two subspecies with different demographic histories. Molecular Ecology, 10(7), 1679-1688.
• Genetic and morphometric differentiation between island and mainland southern elephant seal populations
  
  Hoelzel, A., Campagna, C., & Arnbom, T. (2001). Genetic and morphometric differentiation between island and mainland southern elephant seal populations. Proceedings of the Royal Society B: Biological Sciences, 268(1464), 325-332.
• Molecular genetic diversity and evolution at the MHC DQB locus in four species of pinnipeds
  
  Hoelzel, A., Stephens, J., & O’Brien, S. (1999). Molecular genetic diversity and evolution at the MHC DQB locus in four species of pinnipeds. Molecular Biology and Evolution, 16(5), 611-618.
• Alpha-male paternity in elephant seals
  
  Hoelzel, A., LeBoeuf, B., Reiter, J., & Campagna, C. (1999). Alpha-male paternity in elephant seals. Behavioral Ecology and Sociobiology, 46, 298-306.
• Impact of population bottlenecks on genetic variation and the importance of life-history: a case study of the northern elephant seal
  
  Hoelzel, A. (1999). Impact of population bottlenecks on genetic variation and the importance of life-history: a case study of the northern elephant seal. Biological Journal of the Linnean Society, 68, 23-39. https://doi.org/10.1111/j.1095-8312.1999.tb01156.x
• Molecular population genetics of the southern elephant seal Mirounga leonina
  
  Slade, R., Moritz, C., Hoelzel, A., & Burton, H. (1998). Molecular population genetics of the southern elephant seal Mirounga leonina. Genetics, 149(4), 1945-1957.
• Low genetic variation among killer whales (Orcinus orca) in the eastern North Pacific and genetic differentiation between foraging specialists
  
  Hoelzel, A., Dahlheim, M., & Stern, S. (1998). Low genetic variation among killer whales (Orcinus orca) in the eastern North Pacific and genetic differentiation between foraging specialists. Journal of Heredity, 89(2), 121-128.
• Genetic differentiation between parapatric 'nearshore' and 'offshore' populations of the bottlenose dolphin
  
  Hoelzel, A., Potter, C., & Best, P. (1998). Genetic differentiation between parapatric ’nearshore’ and ’offshore’ populations of the bottlenose dolphin. Proceedings of the Royal Society B: Biological Sciences, 265, 1177-1183.
• Genetic structure of cetacean populations in sympatry, parapatry and mixed assemblages; implications for conservation policy.
  
  Hoelzel, A. (1998). Genetic structure of cetacean populations in sympatry, parapatry and mixed assemblages; implications for conservation policy. Journal of Heredity, 89, 451-458. https://doi.org/10.1093/jhered/89.5.451
• RAPID EVOLUTION OF A HETEROPLASMIC REPETITIVE SEQUENCE IN THE MITOCHONDRIAL-DNA CONTROL REGION OF CARNIVORES
  
  Hoelzel, A., Lopez, J., Dover, G., & Obrien, S. (1994). RAPID EVOLUTION OF A HETEROPLASMIC REPETITIVE SEQUENCE IN THE MITOCHONDRIAL-DNA CONTROL REGION OF CARNIVORES. Journal of Molecular Evolution, 39(2), 191-199.
• IMPROVED RESOLUTION OF (CAC)(N) DNA FINGERPRINTS
  
  Hoelzel, A. (1994). IMPROVED RESOLUTION OF (CAC)(N) DNA FINGERPRINTS. Nucleic Acids Research, 22(7), 1315-1315.
• Ecology and genetics of whales and dolphins
  
  Hoelzel, A. (1994). Ecology and genetics of whales and dolphins. Annual Review of Ecology, Evolution, and Systematics, 25, 377-399. https://doi.org/10.1146/annurev.es.25.110194.002113
• ELEPHANT SEAL GENETIC-VARIATION AND THE USE OF SIMULATION-MODELS TO INVESTIGATE HISTORICAL POPULATION BOTTLENECKS
  
  Hoelzel, A., Halley, J., Obrien, S., Campagna, C., Arnbom, T., Leboeuf, B., Ralls, K., & Dover, G. (1993). ELEPHANT SEAL GENETIC-VARIATION AND THE USE OF SIMULATION-MODELS TO INVESTIGATE HISTORICAL POPULATION BOTTLENECKS. Journal of Heredity, 84(6), 443-449.
• FORAGING BEHAVIOR AND SOCIAL GROUP-DYNAMICS IN PUGET-SOUND KILLER WHALES
  
  Hoelzel, A. (1993). FORAGING BEHAVIOR AND SOCIAL GROUP-DYNAMICS IN PUGET-SOUND KILLER WHALES. Animal Behaviour, 45(3), 581-591. https://doi.org/10.1006/anbe.1993.1068
• Generation of VNTRs and heteroplasmy by sequence turnover in the mitochondrial control region of two elephant seal species.
  
  Hoelzel, A., Hancock, J., & Dover, G. (1993). Generation of VNTRs and heteroplasmy by sequence turnover in the mitochondrial control region of two elephant seal species. Journal of Molecular Evolution, 37, 190-197. https://doi.org/10.1007/bf02407355
• Conservation genetics of whales and dolphins
  
  Hoelzel, A. (1992). Conservation genetics of whales and dolphins. Molecular Ecology, 1(2), 119-125.
• APPLICATIONS OF MOLECULAR GENETIC TECHNIQUES TO THE CONSERVATION OF SMALL POPULATIONS
  
  Amos, B., & Hoelzel, A. (1992). APPLICATIONS OF MOLECULAR GENETIC TECHNIQUES TO THE CONSERVATION OF SMALL POPULATIONS. Biological Conservation, 61(2), 133-144.
• EVOLUTION OF THE CETACEAN MITOCHONDRIAL D-LOOP REGION
  
  Hoelzel, A., Hancock, J., & Dover, G. (1991). EVOLUTION OF THE CETACEAN MITOCHONDRIAL D-LOOP REGION. Molecular Biology and Evolution, 8(4), 475-493.
• GENETIC DIFFERENTIATION BETWEEN SYMPATRIC KILLER WHALE POPULATIONS
  
  Hoelzel, A., & Dover, G. (1991). GENETIC DIFFERENTIATION BETWEEN SYMPATRIC KILLER WHALE POPULATIONS. Heredity, 66, 191-195.
• Whaling in the dark
  
  Hoelzel, A. (1991). Whaling in the dark. Nature, 352(481). https://doi.org/10.1038/352481a0
• A PATERNITY TEST CASE FOR THE KILLER WHALE (ORCINUS-ORCA) BY DNA FINGERPRINTING
  
  Hoelzel, A., Ford, J., & Dover, G. (1991). A PATERNITY TEST CASE FOR THE KILLER WHALE (ORCINUS-ORCA) BY DNA FINGERPRINTING. Marine Mammal Science, 7(1), 35-43.
• KILLER WHALE PREDATION ON MARINE MAMMALS AT PUNTA-NORTE, ARGENTINA - FOOD SHARING, PROVISIONING AND FORAGING STRATEGY
  
  Hoelzel, A. (1991). KILLER WHALE PREDATION ON MARINE MAMMALS AT PUNTA-NORTE, ARGENTINA - FOOD SHARING, PROVISIONING AND FORAGING STRATEGY. Behavioral Ecology and Sociobiology, 29(3), 197-204.
• THE FORAGING SPECIALIZATIONS OF INDIVIDUAL MINKE WHALES
  
  Hoelzel, A., Dorsey, E., & Stern, S. (1989). THE FORAGING SPECIALIZATIONS OF INDIVIDUAL MINKE WHALES. Animal Behaviour, 38, 786-794. https://doi.org/10.1016/s0003-3472%2889%2980111-3
• TERRITORIAL BEHAVIOR OF THE ROBIN ERITHACUS-RUBECULA - THE IMPORTANCE OF VEGETATION DENSITY
  
  Hoelzel, A. (1989). TERRITORIAL BEHAVIOR OF THE ROBIN ERITHACUS-RUBECULA - THE IMPORTANCE OF VEGETATION DENSITY. Ibis, 131(3), 432-436.
• DNA fingerprinting and 'scientific' whaling.
  
  Hoelzel, A., & Amos, W. (1988). DNA fingerprinting and ’scientific’ whaling. Nature, 333(6171). https://doi.org/10.1038/333305a0
• SONG CHARACTERISTICS AND RESPONSE TO PLAYBACK OF MALE AND FEMALE ROBINS ERITHACUS-RUBECULA
  
  Hoelzel, A. (1986). SONG CHARACTERISTICS AND RESPONSE TO PLAYBACK OF MALE AND FEMALE ROBINS ERITHACUS-RUBECULA. Ibis, 128(1), 115-127.